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-Jawless, deep sea scavengers, body fluids are isotonic with sea water. -slime glands for defense |
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-New Synapomorhpy: Vertebral structure with cartilaginous skeletal elements -homologous only with the neural arches of jawed vertebrates -protrusible tongue-like structure |
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JAWS!! -critical step in evolution -now have the ability to cut food and grasp it -Derived features: -two sets of paired fins or limbs -two olfactory tracts -improvements in locomotor/predatory abilities with more complex vertebrae -Ribs (increased anchorage for muscles -Cerebellum ALL MADE POSSIBLE BY SECOND HOW GENE DUPLICATION! |
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key part of gnathostome evolution -jaw was initially important for gill ventilation to respond to increased metabolic needs, helped with forced ventilation of water over gills |
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-Cartilaginous fish -New Synapomorphy: tesserate or prismatic endoskeletal calcification |
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family under Chondrichtheys, have multiple gill openings |
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Squalomorpha and Galeomorpha |
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two families under Neoselachii, two different lineages of sharks. Sq=cold, deep water while Gale=warmer, shallow water |
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Under Neoselachii, Skates have thick tail stalk with 2 dorsal fins and caudal fin while Rays have whiplike tail with barbs |
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Ratfishes, small, soft anatomy, ancient branch of Chondrichtheys |
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Bony Fish! New Synapomorhpyh is calcified bone(Endochondral bone), appeared during Devonian |
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Lineage of Osteichtheys with "lobe fins" that have massive jaw muscles which produced skull changes that separated them -coated with cosmine, a dentine like structure -gave rise to tetrapods |
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(Lungfish)one lineage under Sarcopterygii -weakly developed gill, use lungs - gave rise to tetrapod lungs -fused dorsal, caudal, anal fins with cartilaginous skeleton. -Estivation is the burrowing when waters recede |
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Ray Finned Fishes! -largest radiation -brains everted as formed to give more space -thick, interlocking scales gave more mobility and maneuverability |
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Next synapomorphy after actinopterygii -gars (Lepisosteiformes) and bowfin (Amiiformes) fall under Holosteans -Bony skin -main group is the teleosts -Main SYNAPOMORPHY was a new jaw mechanism where the cheek was no longer solid, allowing for more expansion and a stronger bite (adductor muscles weren't limited by solid cheek) |
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Major Changes -Specialization in jaw to increase suction -protusible premaxilla and maxilla of primary jaw to increase suction -specialized mobile jaws -caudal fin had increased support with Certotrichia spines in fin |
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-Majority of extant teleosts -have sticky eggs and the Weberian apparatus (using gas bladder as tempanum, connects swim bladder to auditory system) |
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marine teleosts, threatened by outflow of water and inflow of salt -low kidney filtration rate with less urine -Drink seawater to maintain water levels while gills pump salt out of body. |
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freshwater -threatened by inflow of water and outflow of salt -gills actively transport salt into body -large amounts of dilute urine with salts reabsorbed |
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has small range of salinity that it can adapt to |
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can live in wide range of salinity |
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fertilized eggs develop outside female (external fertilization) |
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typically associated with Oviparity, the eggs float in water column |
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internal fertilization, but young depend on yolk of the fertilized eggs. |
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Eggs sing to bottom, typically less of them and buried in gravel, float on bottom, or adhere |
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live birth! internal fertilization where young depend on placenta |
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Shared, derived characters -Single Element vertebral column -Bicusped, pedicellate teeth (uncalcified zone at base) -Reduced skull table bones -Moist vascular skin -carnivorous with mucous and poison glands |
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Salamanders -elongate body -simple, primitive limbs which reflect habitat -movement is fish like undulation |
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Frogs -specialized, long hindlimbs for jumping -long toes -pelvic girdle enlarged, strengthened, and anchored to vertebral column -no tail |
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Caecilians -worm like -legless -mostly burrowing -solid skull -up to 200 vertebra |
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Coalescence of Continents into Pangea -extensive shallow seas -land uplifted for freshwater basins -first forests and insects MAJOR OPP. FOR SPECIATION |
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-Main things that distinguish them from invertebrates is: Hox gene complex and the neural crest (the 4th germ layer) |
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