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Unit 3 - New
3rd unit (plants)
178
Biology
Undergraduate 1
04/07/2018

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Term
A vascular plant consists of a
Definition
a root system and a shoot system.
Term
Roots and shoots grow at their
Definition
tips, which are called apices (singular, apex).
Term
The root system
Definition
anchors the plant and penetrates the soil, from which it absorbs water and minerals. These are often extensive, and growing roots can exert great force to move matter as they grow.
Term
The shoot system
Definition
consists of the stems and their leaves.
Term
Roots evolved
Definition
later than shot system as an adaptation to living on land. Flowers, other reproductive organs and fruits and seeds are also formed on the shoot.
Term
Stems serve as a
Definition
scaffold for positioning the leaves, the principle sites for positioning the leaves.
Term
An axillary bud
Definition
is a lateral shoot apex that allows the plant to branch or replace the main shoot if it is eaten by an herbivore.
Term
A vegetative axillary bud
Definition
has the capacity to replace the primary shoot.
Term
Plant cell types can be distinguished by the
Definition
size of their vacuoles, wether they are living or not at maturity, and by the thickness of their cellulose cell walls.
Term
Cells that support the plant are
Definition
more heavily reinforced cell walls with multiple layers of cellulose and other strengthening molecules such as lignin and pectin.
Term
The three basic types of tissues in plants include:
Definition
dermal, ground, and vascular tissue.
Term
Dermal tissue
Definition
primarily epidermis, is one cell layer thick in most plants and forms an outer protective covering for the plant.
Term
Ground tissue
Definition
function in storage, photosynthesis, and secretion, in addition to forming fibers that support and protect plants.
Term
Vascular tissue
Definition
conducts fluids and dissolved substances throughout the plant body.
Term
Meristem cells
Definition
are undifferentiated cells that can divide indefinitely and give rise to many types of differentiated cells. These are clusters of small cells with dense cytoplasm and large nuclei which act as stem cells do in animals.
Term
The extension of both the root and shoot system occurs as a result of repeated cell divisions and subsequent elongation of the cells produced by
Definition
apical meristems.
Term
In woody plants,
Definition
lateral meristems produce an increase in root and shoot diameter.
Term
Apical meristems are located
Definition
at the tips of stems and roots.
Term
The tissues derived from apical meristems are called
Definition
primary tissues
Term
Primary tissues
Definition
are tissues that make up the plant body
Term
Primary plant body
Definition
the extension of the root and stems
Term
The three primary meristems
Definition
protoderm, precambium and ground meristem
Term
Protoderm
Definition
forms the epidermis
Term
Ground meristem
Definition
differentiates further into ground tissue
Term
Procambium
Definition
produces primary vascular tissues
Term
intercalary meristems
Definition
arise in stem internodes (spaces between leaf attachments), adding to the internode lengths.
Term
Lateral meristems
Definition
produce secondary growth. This is the peripheral cylinders of meristematic tissue within the stems and roots that increase the diameter of gymnosperms and most angiosperms.
Term
In woody plants there are two lateral meristems.
Definition
Cork cambium and vascular cambium. The Vascular meristem is just beneath the cork cambium.
Term
Cork cambium
Definition
is a lateral meristem that contributes to the outer bark of a tree
Term
Vascular cambium
Definition
a lateral meristem that produces secondary vascular tissue. It forms between the xylem and phloem in vascular bundles, adding secondary vascular tissues to both sides. Xylem is added to the inside of the vascular cambium and phloem is on the outside.
Term
Secondary tissues
Definition
formed from lateral meristems in trees and shrubs
Term
Secondary plant body
Definition
the part of a plant that consists of secondary tissues from lateral meristem tissues; the older trunk, branches and roots of woody plants.
Term
The epidermis is formed from the
Definition
apical meristem.
Term
The epidermis
Definition
is one cell layer thick in most plants and forms the outer protective covering of the plant. This is a dermal tissue.
Term
In young, exposed parts of the plant, the epidermis is covered with a
Definition
fatty cutin layer constituting the cuticle
Term
Cuticle
Definition
a waxy or fatty noncellular layer (formed of cutin) on the outer wall of the epidermis. Helps to limit water loss and protects from UV damage.
Term
Guard cells
Definition
are paired, sausage shaped cells flanking a stoma. Contains chloroplasts.
Term
Stomata
Definition
a mouth-shaped epidermal opening. Water vapor and carbon dioxide is let in and oxygen is released.
Term
The TMM gene
Definition
encodes a membrane-bound receptor that is part of a signaling pathway controlling asymmetrical cell division.
Term
Trichomes
Definition
are cellular or multicellular hairlike outgrowths of the epidermis. They occur frequently on stems, leaves, and reproductive organs. These reduce evaporation by covering stomatal openings. They also protect leaves from UV radiation and high-intensity lights. Some are glandular, secreting sticky or toxic substances.
Term
Root hairs
Definition
are tubular extensions of individual epidermal cells, and occur in a zone just behind the tips of young, growing roots. No cell wall isolates the hair from the rest of the cell. Increases the root's surface area and efficiency of absorption.
Term
As a root grows
Definition
, the extent of the root hair zone remains roughly constant as root hairs at the older end slough off while new ones are produced at the apex.
Term
Ground tissue consists
Definition
of thin-walled parenchyma cells that function in storage, secretion, and photosynthesis. Other ground tissues, composed of collenchyma cells and sclerenchyma cells provide support and protection.
Term
Parenchyma cells
Definition
are most common type of plant cell. They have large vacuoles, thin walls, and are initially (but briefly) more or less spherical. These cells, which have living protoplasts, push ip against each other shortly after they are produced, however, and assume other shapes and often end up with 11 to 17 sides. They may live for many years and function in storage of food and water, photosynthesis and secretion.
Term
Some parenchyma cells contain
Definition
chloroplasts, especially in leaves and the outer parts of herbaceous stems.
Term
Collenchyma cells, like parenchyma cells,
Definition
have living protoplasts, lack cell walls and may live for many years. These cells are usually a little longer than wide, have walls that vary in thickness. These flexible cells provide support for plant organs, allowing them to bend without breaking. They often form strands or continuous cylinders beneath the epidermis of cells or leaf stalks and along veins in leaves.
Term
Sclerenchyma cells
Definition
have thick walls. They are usually dead at maturity. Their secondary cell walls often contain lignin. Occurs in two types; fivers and sclereids.
Term
Lignin
Definition
is a highly branched polymer that makes cell walls more rigid.
Term
Fibers are
Definition
long and slender cells that are usually grouped together in strands.
Term
sclereids are
Definition
variable in shape but often branched. May occur singly or in groups; aren't elongated, but may have different forms, including that of a star.
Term
Vascular tissue includes two types of conducting tissues;
Definition
xylem and phloem.
Term
Xylem
Definition
conducts water and dissolved minerals. Contains a combination of vessels. Also supports plant body. Includes fibers and parenchyma cells.
Term
Phloem
Definition
conducts a solution of carbohydrates (mainly sucrose). Also transports hormones, amino acids, and other substances needed for cell growth. Located on the outer part of roots and stems. Two kinds of elongated cells; sieve cells and sieve-tube mebers.
Term
Vessels
Definition
continuous tubes formed from dead, hollow, cylindrical cells arranged end-to-end.
Term
Tracheids
Definition
dead cells that taper at the ends and overlap one another. Contains pits
Term
Transpiration
Definition
The loss of water vapor by plant parts; mainly in stomata
Term
Vessel members tend to be
Definition
shorter and wider than tracheids.
Term
Both vessel memvers and tracheids have
Definition
thick, lignified secondary walls and no living protoplasts at maturity.
Term
Pits are
Definition
small areas where no secondary wall material has been deposited. The primary wall is present but it is water permeable. There is a continuous flow of water from tracheid to tracheid
Term
Sieve cells
Definition
less specialized than sieve-tube members, and the pores in all of their sieve areas are about the same diameter.
Term
Sieve-tube members
Definition
Stacked up end to end in the phloem forming longitudinal series called sieve tubes. More efficient than sieve cells.
Term
Each sieve tube member is associated with an adjacent, parenchyma cell known as a companion cells. The companion cells
Definition
carry our some of the metabolic functions needed to maintain the associated sieve-tube member, because the sieve tube member lacks a nucleus.
Term
There are 4 regions in developing roots
Definition
the root cap, the zone of cell division, the zone of elongation and the zone of maturation
Term
The root cap
Definition
is composed of two types of cells, inner columella cells and the outer lateral root cap cells. Protects the delicate tissues behind it as it moves through the soil.
Term
Columella cells contain
Definition
amyloplasts (plastids with starch grains) that collect on the sideds of cells facing the pull of gravity.
Term
The zone of cell division
Definition
most of the activity takes place toward the edges of the meristem, which is located in the center of the root tip and protected by the root cap, where the cells divide every 12 to 37 hours.
Term
A group of cells in the center of the root apical meristem
Definition
called the quiescent center, divides only very infrequently.
Term
The zone of elongation
Definition
is where roots lengthen because the cells produced by the primary meristems elongate. The small vacuoles merge and grow till they occupy 90% or more of the volume of each cell. Cells don't increase past the zone of elongation.
Term
The zone of maturation
Definition
The cells that have elongated in the zone of elongation become differentiated into specific cell types
Term
The cortex
Definition
may be many cell layers wide and functions in food storage
Term
Casparian strips
Definition
surround each adjacent endodermal cell wall perpendicular to the root's surface. Blocks transport between cells.
Term
The Stele
Definition
The center vascular system of stems and roots
Term
Pericycle
Definition
one or more cell layers that surrounds the vascular tissues of the root, bounded externally by the endodermis and internally by the phloem.
Term
Adventitious root
Definition
any root that arises along a stem or in some place other than the root
Term
Prop roots
Definition
These so-called prop roots grow down to the ground and brace the plants against wind
Term
aerial roots
Definition
which are attached to tree branches and grow unconnected to the ground (but are not parasites), have roots that extend into the air. Some aerial roots have an epidermis that is several cell layers thick, an adaptation to reduce water loss. These aerial roots may also be green and photosynthetic
Term
Pnumatophores
Definition
Some plants that grow in swamps and other wet places may produce spongy outgrowths called pneumatophores from their underwater roots. The pneumatophores commonly extend several centimeters above water, facilitating oxygen uptake in the roots beneath
Term
Parasitic Roots
Definition
penetrate the host plants around which they are twined. The haustoria establish contact with the conducting tissues of the host and effectively parasitize their host. Dodder not only weakens plants but can also spread disease when it grows and attaches to several plants.
Term
Food storage roots
Definition
produce extra parenchyma cells that store large quantities of carbohydrates. Carrots, beets, parsnips, radishes, and turnips have combinations of stem and root tissues that also function in food storage. Cross sections of these roots reveal multiple rings of secondary growth.
Term
Water storage roots
Definition
arid regions, may produce water storage roots weighing 50 kg or more
Term
Buttress roots
Definition
produce huge buttress roots toward the base of the trunk, which provide considerable stability
Term
Contractile roots
Definition
contract by spiraling to pull the plant a little deeper into the soil each year, until they reach an area of relatively stable temperature.
Term
Taproot system
Definition
Single large root with small branch roots
Term
Fibrous root system
Definition
many small roots of similar diameter
Term
Node
Definition
point of attachment of leaf to stem
Term
Internode
Definition
area of stem between two nodes
Term
Blade
Definition
flattened part of leaf
Term
Petiole
Definition
stalk of leaf
Term
Axil
Definition
angle between petiole/blade and stem
Term
Axilliary Bud
Definition
develops into branches with leaves or may form flowers
Term
Terminal Bud
Definition
extends the shoot system during the growing season
Term
Lenticels
Definition
Cork cambium produces unsuberized cells that permit gas exchange to continue
Term
Runners and stolons
Definition
horizontal stems with long internodes that grow along the surface of the ground
Term
Microphyll
Definition
Leaf with one vein branching from the vascular cylinder of the stem and not extending the full length of the leaf
Term
Megaphylls
Definition
Several to many veins
(most plants)
Term
Eudicot Leaves
Definition
Most eudicot leaves have a flattened petiole
Slender stalk called petiole
Leaf flattening increases photosynthetic surface
Flattening of the leaf blade reflects a shift from radial symmetry to dorsal–ventral (top–bottom) symmetry
Term
stipules
Definition
Outgrowths at base of petiole
May be leaf-forming or modified as spines
Term
Veins
Definition
Vascular bundles in leaves
Main veins are parallel in most monocot leaves
Veins of eudicots form an often intricate network
Term
Simple leaves
Definition
contain undivided blades
May have teeth, indentations, or lobes
Term
Compound leaves have
Definition
blades that are divided into leaflets
Term
Most eudicot leaves have 2 types of mesophyll
Definition
Palisade mesophyll and Spongy mesophyll
Term
Spongy mesophyll
Definition
loosely arranged cells with many air spaces in between
Term
Palisade mesophyll
Definition
usually two rows of tightly packed chlorenchyma cells
Term
Monocot leaves
Definition
mesophyll is usually not differentiated into palisade/spongy layers
Term
Floral leaves (bracts)
Definition
surround true flowers and behave as showy petals
Term
Spines
Definition
reduce water loss and may deter predators
Term
Reproductive leaves
Definition
plantlets capable of growing independently into full-sized plant
Term
Window leaves
Definition
succulent, cone-shaped leaves that allow photosynthesis underground
Term
Shade leaves
Definition
larger in surface area but with less mesophyll than sun-lit leaves
Term
Insectivorous leaves
Definition
trap insects
Term
Aquaporins
Definition
water-selective pores in plasma membrane that increase the rate of osmosis because they allow bulk flow
Term
Osmosis
Definition
If a single plant cell is placed into water
Water moves into cell by osmosis
Cell expands and becomes turgid
If cell placed in high concentration of sucrose
Water leaves cell
Cell shrinks – plasmolysis
Term
Apoplast route
Definition
movement through the cell walls and the space between cells
Avoids membrane transport
Term
Symplast route
Definition
cytoplasm continuum between cells connected by plasmodesmata
Term
Transmembrane route
Definition
membrane transport between cells and across the membranes of vacuoles within cells
Permits the greatest control
Term
Phloem-loading
Definition
occurs at the source
Carbohydrates enter the sieve tubes in the smallest veins at the source
Sieve cells must be alive to use active transport to load sucrose
Water flows into sieve tubes by osmosis
Turgor pressure drives fluid throughout plant
At sink, sucrose actively removed and water follows by osmosis
Water may be recirculated in xylem or lost
Term
Pressure-flow theory
Definition
is a model describing the movement of carbohydrates in phloem
Dissolved carbohydrates flow from a source and are released at a sink
Sources include photosynthetic tissues
Food-storage tissue can be sources or sinks
Sinks include growing root and stem tips as well as developing fruits
Term
Halophytes
Definition
Plants that can tolerate soils with high salt concentrations
Term
Over 90% of the water taken in by the plant’s roots is ultimately
Definition
lost to the atmosphere
Term
Cavitation
Definition
An air bubble can break the tensile strength of a water column
A gas-filled bubble can expand and block the tracheid or vessel
Term
Soil
Definition
Highly weathered outer layer of the Earth’s crust
Mixture of sand, rocks, clay, silt, humus, and mineral and organic matter
The Earth’s crust includes about 92 naturally occurring elements
Most are found in the form of inorganic compounds called minerals
Also full of microorganisms
Term
Topsoil
Definition
Most roots are found in topsoil
Mixture of mineral particles of varying sizes, living organisms, and humus
Characterized by their relative amounts of sand, silt, and clay
Soil composition determines the degree of water and nutrient binding to soil particles
Term
Only minerals dissolved in water in spaces among soil particles are
Definition
available for uptake by roots
Term
Pores in Soil
Definition
About half of the soil volume is occupied by pores
May be filled with air or water
Term
If topsoil is lost
Definition
soil’s water-holding capacity and nutrient content are adversely affected
Term
Measures to prevent erosion
Definition
Intercropping
Conservation tillage
No-till
Term
Measures to prevent fertilizer runoff
Definition
Site-specific farming
Integrated nutrient management
Term
Acidic soils release minerals, such as aluminum
Definition
that are toxic to plants
Term
Saline soils alter water potential, leading to a
Definition
loss of water and turgor in plants
Term
Macronutrients -used in relatively large amounts
- needed in plants
Definition
Nine = C, O, H, N, K, Ca, Mg, P, and S
Term
Micronutrients – used in minute amounts - needed in plants
Definition
Seven = Cl, Fe, Mn, Zn, B, Cu, and Mo
Term
Photosynthesis is major source of
Definition
plant nutrition via the fixation of _CO_2_ into sugar using solar energy
Term
Hydroponic Culture
Definition
plant roots are suspended in aerated water containing nutrients

Plant seedling is first grown in a complete nutrient solution
Seedling is then transplanted to a solution lacking one suspected essential nutrient
Growth of the seedling is monitored for presence of abnormal symptoms
Term
Plants need ammonia _(NH_3_) or nitrate _(NO_3_−_) to ______

However, they lack the biochemical pathways necessary to convert _____
Definition
to build amino acids

_N_2_ to _NH_3_
Term
Rhizobium bacteria require oxygen and carbohydrates to support their energetically expensive lifestyle as nitrogen fixers
Definition
Plant host supplies both
Term
Symbiotic associations with mycorrhizal fungi are found in about
Definition
90% of vascular plants
Substantially expand the surface area available for nutrient uptake
Enhance uptake of phosphorus and micronutrients
Term
Carnivorous Plants
Definition
Often grow in acidic soils that lack nitrogen
Trap and digest small animals, primarily insects, to extract additional nutrients
Have modified leaves adapted for luring and trapping prey
Prey is digested with enzymes secreted from specialized glands
Term
Pitcher plants
Definition
Have pitcher-shaped leaves with cavity filled with digestive fluid
Term
Venus flytrap
Definition
When hairs are touched, the two halves of the leaf snap together
Term
Sundews
Definition
Glandular trichomes secrete both sticky mucilage and digestive enzymes
Term
Waterwheel
Definition
Uses trigger hairs and snaps to capture and digest small aquatic animals
Term
The snap-trap mechanism was acquired by a common ancestor of
Definition
the Venus flytrap and the aquatic waterwheel
Term
Dodder
Definition
(nonphotosynthetic)
Wraps around its host
Relies on host for its nutritional needs
Term
Indian pipe
Definition
(nonphotosynthetic)
Hooks into host trees through mycorrhizae
Term
Calvin cycle fixes
Definition
_CO_2_ into sugar
Ribulose 1,5-bisphosphate carboxylase/oxygenase (rubisco) catalyzes the first step
Can bind _CO_2_ or _O_2_
If _CO_2_ binds, a 3-C sugar is made, that can be used to make glucose and sucrose
If _O_2_ binds, photorespiration occurs
Neither nutrient nor energy storage
Term
_C_3_ photosynthesis occurs in
Definition
mesophyll cells
Term
_C_4_ photosynthesis uses
Definition
an extra pathway to shuttle carbon deep within the leaf
This reduces photorespiration by limiting the Calvin cycle to cells surrounding the vascular tissue where _O_2_ levels are low
Term
In _C_3_ plants, as _CO_2_ increases
Definition
the Calvin cycle becomes more efficient
Term
As _CO_2_ levels increase,
Definition
relatively less nitrogen and other macronutrients are found in leaves
Term
Phytoremediation
Definition
Use of plants to concentrate or breakdown pollutants
Term
Phytodegradation
Definition
contaminant is taken up from soil and broken down
Term
Phytovolatilization
Definition
contaminant is taken up from soil and released through stomata
Term
Phytoaccumulation
Definition
contaminant is taken up from soil and concentrated in shoots
These are later harvested
Term
Trichloroethylene (TCE)
Definition
May be removed from the soil by poplar trees
Degraded into _CO_2_ and chlorine
A fraction moves rapidly through the xylem and is released through stomata
Term
Trinitrotoluene (TNT)
Definition
May be removed from soil and degraded by poplar and bean plants
But at high concentrations, it is toxic to these plants
Term
Heavy metals, including arsenic, cadmium, and lead, are
Definition
toxic to animals in even small quantities
Term
400 plant species have the ability to
Definition
hyperaccumulate toxic metals from soi;
Term
Invasive Species
Definition
One of the greatest problems with nonnative invasive species, such as the emerald ash borer, is the lack of natural predators in the new environment
Term
Dermal Tissue System
Definition
First-line defense of all plants
Epidermal cells throughout the plant secrete wax to protect plant surfaces from water loss and attack
Above-ground parts also covered with cutin
Suberin is found in cell walls of subterranean plant organs
Silica inclusions, trichomes, bark, and even thorns can also offer protection
Term
Mechanical wounds allow
Definition
microbial entry
Term
Phases of fungal invasion
Definition
Windblown spore lands on leaves
Spore germinates and forms adhesion pad
Hyphae grow through cell walls and press against cell membrane
Hyphae differentiate into haustoria
Term
Fungi and bacteria can also be beneficial to plants
Definition
Mycorrhizal fungi
Nitrogen-fixing bacteria like Rhizobium
Plant growth-promoting rhizobia (PGPR)

Bacteria provide substances that support plant growth
Can also limit the growth of pathogenic soil bacteria
Term
Many plants produce toxins that
Definition
kill herbivores, make them ill
Term
Metabolic pathways needed to
Definition
sustain life are modified
Term
Plants protect themselves from toxins in two main ways
Definition
Sequester a toxin in a membrane-bound structure

Produce a compound that is not toxic until it is metabolized by attacking animal
Cyanogenic glycosides break down into cyanide (HCN) when ingested
Term
Allelopathic Plants
Definition
Secrete chemicals to block seed germination or inhibit growth of nearby plants
This strategy minimizes competition for resources
Term
Ricin
Definition
Ricin is an alkaloid produced by the castor bean plant (Ricinus communis)
It is six times more lethal than cyanide and twice as lethal as cobra venom
A single seed can kill a small child
It functions as a ribosome-binding protein that inhibits translation
Term
Many secondary metabolites have
Definition
benefits to human health
Term
Phytoestrogens of soy plants
Definition
Appear to lower the rate of prostate cancer in Asian males
However, questions have been raised about their effect on unborn babies
Also on babies consuming soy-based formula
Term
Taxol of Pacific yew trees
Definition
Fights cancers, especially breast cancer
Term
Quinine of Cinchona trees
Definition
Effective against malaria, which is caused by four species of Plasmodium
Blocks DNA replication
Also leads to build-up of toxic hemes that poison the parasite
Term
Complex coevolution of plants and animals has resulted in
Definition
mutualistic associations
Term
Acacia trees and ants
Definition
Small armies of ants protect Acacia trees from harmful herbivores
Plant provides ants with food and shelter
Term
Wound Response Signaling
Definition
Wounded leaves produce an 18-amino acid peptide called systemin
Systemin moves throughout the plant in the phloem
Cells with receptors produce jasmonic acid
Jasmonic acid turns on genes for proteinase inhibitor
Term
H. H. Flor’s gene-for-gene hypothesis
Definition
Plants have a plant resistance gene (R); pathogens have an avirulence gene (avr)
It is the recognition of the gene products (i.e. proteins) that is critical
If binding occurs, plant can mount defenses that keep pathogen avirulent
If no binding occurs, the plant succumbs to disease
Term
Hypersensitive Response
Definition
Recognition of the pathogen by the R gene product leads to hypersensitive response
Leads to a very rapid cell death around the site of attack
Also to longer term, whole plant resistance
Term
Rapid Cell Death
Definition
Rapid cell death due to hypersensitive response
Seals off the wounded tissue to prevent the pathogen or pest from moving into rest of the plant
Hydrogen peroxide and nitric oxide produced
May signal cascade of chemical events resulting in localized host cell death
Phytoalexins – antimicrobial chemical defense agents
Term
Systemic Acquired Resistance (SAR)
Definition
systemic response by plants
Several pathways lead to broad-ranging resistance that lasts for a period of days
Long-distance inducer is likely salicylic acid
At the cellular level, jasmonic acid is involved in SAR signaling
SAR allows the plant to respond more quickly to a second attack
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