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“Study of factors that determine the distribution and abundance of organisms” also: study of interactions of organisms with their environments |
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Finnish ecologist, did research on caterpillars and nitrogen-enriched soils. Lab and field showed 2 different things. Field was affected by predators. |
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How to deal with ecological complexity? |
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Seek simplicity and mistrust it. Multiple working hypotheses. Attempts at prediction -> useful postdiction -> “Ecological Forecasting” |
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Observation, Experiments, Models |
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The only true way of obtaining new information, good natural history, consistent long term monitoring, advance mapping and sensing technology |
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- Field or laboratory, replicated manipulated treatments with controls – Whole ecosystem experiments |
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Experimental tools for dealing with ecological variability: |
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1. Replicates 2. Controls 3. Statistics |
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separate (independent) units of study that are treated identically by ecologists, in order to assess variability that arises from factors we didn’t manipulate |
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units that did not experience the focal manipulation. They provide a baseline for comparison, an understanding of how organisms or systems will change over time or space, independent of experimental treatments. |
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tools help us distinguish signal from noise, e.g., test whether differences between treatments (e.g., control vs experimental) are greater than differences within treatments |
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– verbal or mathematical simplifications of reality, intended to capture key processes driving system change over time – Hypotheses: suggested explanations, subject to test(falsifiable) |
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Big Ecology Questions (3) |
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1. What factors determine distribution and abundance of organisms ? 2. What regulates populations ? (next week) 3. How will organisms, ecosystems or the planet respond to changes in environment (climate, land use, extinctions, invasions of new species) ? |
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X Axis: Precipitation Y Axis: Tempurature Yellow: Desert Lime: Grassland Teal: Temperate Forest Green: Tropical Forest Blue: Coniferous Forest White: Artic and Alpine Tundra |
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What causes spatial and temporal variation in temperature and moisture? |
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Same photon flux spread over larger land surface near poles than at equator |
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Tilt of Earth’s axes sets up seasonality of northern and southern hemispheres |
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rising air experiences lower atmospheric pressure, expands in volume, losing temperature. (First Law of Thermodynamics*—work done to expand air parcels comes at expense of heat energy.) If it cools below dew point, air will lose its moisture (as clouds, or precipitation) |
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temperature to which the air would have to cool (at constant pressure and constant water vapor content) in order to reach saturation, at which the air is holding the maximum possible amount of water vapor possible at the existing temperature and pressure. If the air cools below dew point, moisture must be removed from the air. This is accomplished through condensation. This process results in the formation of fog, frost, clouds, or precipitation. |
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Earth is a sphere, not a cylinder. An object at equator is moving east at 24,000 miles per day. If it moves north, the earth beneath moves more slowly, so it veers right. If it moves from north towards equator, also goes right. Reverse is true in southern hemisphere. |
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Earth rotates in which direction |
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most clearly separated by Temperature and Percipitation |
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stratum of rapid temperature change. Can separate oxygenated from hypoxic habitat. |
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Eutropic/Mesotropic/Oligotropic |
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Eutrophic (river, lake estuary): nutrient rich, likely to produce noxious or harmful algal blooms (cyanobacteria, toxic dinoflagellates) Mesotrophic – intermediate nutrient concentrations Oligotrophic – low nutrient concentrations,very clear water (“good” water quality for humans and fish) |
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the largest niche in which a species could persist in the absence of adverse interactions with other species |
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Range of conditions, resource levels, and densities of other species within which an organism or species can survive and reproduce(persist over time). |
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the (generally smaller) niche volume occupied by a species in the presence of interspecific interactions |
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rely on external sources of heat to regulate temperature |
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use their own metabolic heat production to regulate their body temperature. |
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maintaining an internal state with a narrower (and physiologically more favorable) range of conditions than the organism experiences in the external environment |
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Residual population left over from time when environment could support its survival and reproduction, which can no longer replace itself locally |
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: Habitat where death rates exceed birth rates, and organisms are present only because of immigration from Source Habitats (where births exceed deaths) |
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: Resources produced in one habitat that support consumers in a second habitat. |
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: the study of periodic plant and animal life cycle events that are influenced by environmental changes, especially seasonal variations in temperature and precipitation driven by weather and climate. |
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Acclimatization (“acclimation” in the lab) |
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: shifts in the response of an organism to a condition caused by the regime it has experienced in the past. – e.g. trees can tolerate lower temperatures in October than they can if surprised in mid-summer, because they’ve induced new types of proteins and restructured cell membrane phospholipids. |
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: whether the young of the year can grow large enough to survive their first winter. |
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1. Adaptation 2. Exaptation 3. Phylogenetic constraint |
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: trait that confers a performance advantage relative to its absence in an ancestor |
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: confers an advantage, but originated BEFORE the need for it arose |
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(historical baggage) : trait evolved in the past that may not be suited to current environment |
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organism’s lifetime pattern of growth, differentiation, storage, reproduction |
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: sequence of stages through which organism passes to develop from zygote to a reproductive adult producing more zygotes |
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: individual reproducing once per lifetime (salmon, bamboo, Century plant) |
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: individual that reproduces more than once in its lifetime (humans, oak trees, lobsters) |
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develop from zygote to adult with determinant form |
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grow by repeated interations of its parts (modules) into an adult of indeterminate form (coral, poison oak) |
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20 Currencies: energy, nutrients, time
Allocation? |
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growth activity maintenance Reproduction: offspring quality, offspring quantity |
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Life history trade offs of plants or animals |
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• Starting growth early in the season entails risk of freezing • If reproductive resources allocated to larger seeds, plant makes fewer, so incurs more predation risk, and risk of bad luck (unfavorable microsites) • Dispersal: reduces competition with parent, but increases risk of unsuitable habitat. |
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: study of how the vital rate of individuals (birth, death, growth) affect structure and dynamics of populations |
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: group of potentially interbreeding individuals (same species, co-occur in time and space)
based on – density: number of individuals per area or volume – size structure – age distribution – sex ratio… |
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: pattern of distribution of individuals in space (clumped, even, or random = every site has an equal probability of being occupied by an individual, independent of locations of other individuals) |
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Intraspecific vs. Interspecific |
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Intraspecific interactions (between individuals of the same species) vs Interspecific interactions (between individuals of different species) |
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Life (and reproductive) table |
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—summary of age (or size) specific rates of survival and fecundity (progeny per individual) |
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: a group of individuals of the same age, from birth until they all die (or by other methods that approximate this ideal approach) . |
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Population growth/decline equation |
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dN/dt = b Nt – d Nt = (b – d) Nt = r Nt (closed population) r = b – d = per capita rate of population growth (time -1 ) |
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dN/dt = rN [(K-N)/K]
Carrying Capacity K limits. |
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K depends on both the environment and the organism in question |
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r-selected traits – Short life span – Small size – High predator vulnerability – Weak competitor – Good disperser – Many small offspring – Early reproduction |
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K-selected traits – Long life span – Large size – Low vulnerability to predators – Strong competitor – Slower disperser – Fewer but better provisioned offspring – Late reproduction |
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: Estimate of land and water area needed to produce all resources a nation consumes, and to absorb all the waste it generates |
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• Competition (interspecific) (-, -) • Predation, herbivory, parasitism (+, -) • Mutualism (+, +) • Amensalism (0, -) • Commensalism (0, +) |
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Exploitative Apparent Interference |
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: two species compete for a resource that is limiting: in short supply relative to their needs indirect
Image V |
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: two species directly harm each other by toxic allelochemicals, injury or wasting time, increasing risk, etc. direct
Image --- |
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: two species decrease in each other’s presence because they support the increased abundance or vigor of a common predator indirect
Image A |
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is a type of interaction that is beneficial to both species involved. • It may be obligate or facultative. • It may be symbiotic or non-symbiotic. |
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: a group of species that co-occur in time and space |
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depict feeding (or other significant (e.g. population limiting or regulating)) relationships among members of a community |
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: group of entities united by interaction or interdependence to form or act as an entire unit |
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Odd number of tropic levels |
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: green (plants resource limited = “bottom up” limitation) |
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Even number of tropic levels |
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: barren (plants consumer limited = “top down” limitation) |
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: a species that consumes and limits the population of another species that would otherwise dominate the system |
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: A discrete event that frees resources and opens up habitat by killing or removing organisms |
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Characteristics of early vs late successional species: |
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Early: good dispersers, fast growing, allocation to growth and progeny (r-selected), not defense or competitive ability. Tolerance of harsh conditions or abrasion. Late: poorer dispersers, slower growing but better competitors and/or better defended. (K selected) |
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Effect of early species on later species? |
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• Positive (Facilitation)? • Negative (Inhibition)? • Neutral (Tolerance)? |
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: sequential changes, following disturbance, in the dominant biota occupying and affecting a site |
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: on truly bare (sterilized) space--volcanic lava field; rock exfoliation, receding glaciers |
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: space has been opened and de-populated, but some residual biota has survived (more common than primary succession) |
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(e.g. in rotten log or infested acorn) = habitat modified by inhabitants, disappears over time |
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: late successional stage in which dominant species can replace themselves; young of climax species can successfully recruit under established adults |
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—a small fraction of solar radiation is fixed by photosynthetic organisms, but once energy is dissipated as heat, it is never recovered by ecosystem (except as passive energy savings for transport—e.g. ocean or air currents driven by heat and used by |
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—atoms (H, O, C, N, Ca, P ) created in stars, used again and again, end up in different pools, fluxes transfer atoms between pools. |
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: if system is in equilibrium so input (vol time-1) = output (vol time-1) = q, residence time or turnover time, T = Volume/q |
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: downstream transport with periodic cycling by local biology. Retentive ecosystems with short spiral lengths are more efficient- - more biotic production per nutrient flux downstream. |
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: organic production/nutrient flux (mass time-t / mass time-t ) Forests more efficient at producing wood from nutrients if these are retained. Oligotrophic lakes produce more fish per nutrient flux than eutrophic lakes |
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Equilibrium Theory of Island Biogeography” |
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S reached when Immigration + Evolution = Extinction + Emigration, or, if Evolution and Emigration small, when Immigration = Extinction Effects of island size and distance only factors considered. |
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[image] What is at 60 degrees? 30 Degrees? Poles? Equator? |
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30 Degrees Deserts 60 Degrees Tropics Poles cold Equator Hot |
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