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Evolutionary process in which an adaptation in one species leads to the evolution of an adaptation in a species with which it interacts. Also known as Reciprocal adaptation. Can lead to an evolutionary "arms race" |
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In reference to an evolutionary system, continuing adaptation is needed in order for a species to maintain its relative fitness amongst the systems being co-evolved with. (Taken as an allusion to the Red Queen Race from Lewis Carroll's Through the Looking Glass) |
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There are many different ways in which species interact with one another Antagonistic interactions +/- Mutualistic +/+ Competitive interactions -/- Commensalism +/o Amensalism o/- |
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Predator-prey interactions |
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Used to escape or repel predators |
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Toxic prey advertise with warning coloration |
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Non-toxic organisms mimic toxic ones |
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Many toxic organisms converge on common color patterns. |
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Prey species can escape predation many ways - Crypsis (camouflage) - Homotypy – looking like an inedible object - Stop moving - Run away - Develop shells, tough skin, spines, or hair |
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Specialists that dine on one or a few plant species. (Herbivore) |
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Generalists that feed on many different plant species (Herbivore) |
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Secondary metabolites, plant defense against herbivores |
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Thorns and spines, plant defense against herbivores. |
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Reciprocal Adaptations In Plants and Herbivores |
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Plants evolve new secondary metabolite Plants with new chemical selected for Adaptive radiation among plants with the chemical Herbivores evolve resistance to the chemical, and some may even be able to use it as predator defense |
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Host interactions. Bacteria, viruses, and protists |
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Ability to cause disease Pathogen hosts can evolve resistance. Susceptible, infected, recovered (immune) |
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Many have mechanisms for attaching to their hosts. (Lice, leeches, mosquitoes, etc) |
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Plants and Pollinators. To be effective, the system must have: - An attractant or reward that entices a pollinator - Behavior that ensures the pollinator will visit multiple plants - Anatomical features that ensure pollen transport |
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Frugivores Method of seed dispersal |
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Animals that feed on fruit Transport seeds and deposit them with feces so they are automatically fertilized - Not always in favorable locations Fruit ripening coincides with seed maturation so dispersed seeds are ready for planting |
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Some exchange food for care Some exchange food for transport Some exchange services for housing Some exchange services for defense |
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Intraspecific Competition |
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Competition among individuals of the same species. |
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Interspecific Competition |
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Competition among individuals of different species |
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occurs when one competitor uses limited resources more efficiently and eventually eliminates the other competitor |
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the division of resources by coexisting species such that the niche of each species differs by one or more significant factors |
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individuals actively interfere with another’s access to resources - May restrict habitat use |
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individuals reduce the quantities of their shared resources - May lead to competitive exclusion - May lead to coexistence |
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groups of species that exploit the same resource, but in different ways |
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the tendency for characters to be more divergent in sympatric populations than in allopatric populations of the same two species |
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species may use the same resource in the same location, but at different times Predators can influence competition if they prefer one prey species over another Competition can determine species’ niche space |
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Study of the distribution, abundance, demography, and interactions between coexisting populations |
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The number of species in a given community. Determined in part by energy flow |
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Convert light or chemical energy into organic molecules. |
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Indicate the direction of energy flow. Primary producers – plants Primary consumers – herbivores Secondary consumers – predators that feed on herbivores Detritivores – feed on decomposing organic matter Omnivores – feed on multiple trophic levels |
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Measures the rate of energy accumulation |
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Use diverse sources of energy |
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Interconnected food chains that illustrate energy flow through a community |
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Second law of thermodynamics states that every transfer of energy adds to the entropy of the universe Fewer individuals can be supported at higher trophic levels Energy transfers are inefficient Approximately 10% of the energy at one trophic level will be transferred to the next. |
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Species richness is linked to productivity Low energy environments cannot support many species High energy environments can support many species up to a point Highest energy environments have intense competition, which can lead to competitive exclusion |
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The indirect effects that addition or removal of species has on other trophic levels - Predators - Herbivores |
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a species that has a disproportionate effect on its environment relative to its biomass |
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diversity within a single community |
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the change in species composition from one community to another |
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the regional diversity found over a range of communities in a geographic region |
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There is a general trend of decreasing species diversity as latitude increases - time - spatial heterogeneity - specialization - predation |
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Species area relationship |
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the relationship between the size of an area and the number of species it supports Large areas tend to support more species than small areas |
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Based on immigration and extinction rates Equilibrium number of species (S) is influenced by two main factors: - Island size - Distance from the mainland |
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The species composition of a community changes over time. Progress depends on the activity of successful colonists. |
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Occurs in an environment where new substrate is deposited |
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occurs in an environment where disturbance has cleared substrate that previously supported vegetation |
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The final stage of succession |
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a colonizer modifies the environment in a way that facilitates colonization by other species |
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early colonists prevent colonization by other species |
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the perpetuation of the climax community is dependent on periodic disturbance (e.g., lodgepole pine and fire) |
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succession in detritus based communities Takes place without the inclusion of plants (e.g., benthic zones, corpse decay) |
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Species rich communities tend to be more stable More efficient use of resources Natural communities are often more stable than managed communities (e.g., monocultures in agriculture) |
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