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ray finned fishes -scales ganoid, cycloid, or ctenoid (modified cycloid scale)--and placoid -sister taxon of the Sarcopterygians -the old days: w/ Acanthodians, forms the Teleostorni (old name with 3 clades of bony vertebrates) -3 subclasses: Cladistia, Chondrostei, Neopterygii |
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rubbery, thick, lots of protection (gar) |
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modified cycloid scale (very rough, little teeth) |
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Order Polypteri -Bichirs, Ropefishes (rhombic ganoid scales) -Polypteras (Nile River) & Erpetoichthys (ropefish): rubbery, diamond shaped scales |
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first bony, then turn cartilaginous later -many extinct, only extant Order is Acipenseriformes |
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*Heterocercal tail, Intestine with spiral valve, fin rays more numerous than their basals Acipenseridae- Sturgeons Polyodontidae- Paddlefishes (Chinese paddlefish-Psephurus gladius) |
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fin rays equal in #s to their supports in dorsal and anal fins (tight control on raising/lowering fins) |
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bones in a fish that support the dorsal fin -defense, swim, wedge into rocks, sexual advantage |
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long-nosed gar -ganoid scales -breathes at the surface -may enter salt water -up to 3 meters |
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Bowfin (Amia calva) -freshwater, up to 3 feet (20 pounds) -ganoid scales, surface breathing |
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96% of extant fishes -27 anatomical synapomorphies, no molecular ones -most important are the caudal skeleton |
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Subdivision Osteoglossomorpha |
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-Mooneye, Goldeye of Hiodontiformes -Osteoglossiformes (very primitive) bony tongue |
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bony tongue -Pantodon -Knife fish -elephant fishes -Pirarucu (Arapaima gigas) -Arowana |
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swimbladder not connected with ear Leptocephalus larvae (small head as larva) 3 orders: Elapiformes (tarpons), Albuliformes (bonefishes), Anguilliformes (eels & morays) |
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Subdivision Ostarioclupeomorpha |
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Superorder Clupeomorpha Superoder Ostariophysi -inner ear and swim bladder are anatomical synapomorphies -mostly molecular synapomorphies |
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anterior extension of swimbladder enters the skull -clupeiformes (herring) -school of anchovies |
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synapomorphy: fright substance- Schrekstoff hormone Series Anotophysi (milkfish- Gonorynchiformes) Series Otophysi (piranha/catfish) |
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synapomorphy: Weberian apparatus -anatomical structure that connects the swim bladder to the auditory system |
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synapomorphy Retractor dorsalis: RAB |
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large paired muscle that originates along the ventral margin of the anteriormost three or four vertebral centra and inserts on the dorsal-most parts of the gill arches. The muscle functions to retract or pull back on the upper pharyngeal teeth as part of a chewing mechanism that prepares food for swallowing |
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