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Exam 3
Animals
313
Biology
Undergraduate 1
03/26/2012

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Term
Nervous system of Insects
Definition
brain has several subdivisions with separate functions
Term
With increasing complexity, overall trend toward cephalization
Definition
increasingly complex brain in anterior region
Term
vertebrate nervous system
Definition
Central Nervous System (CNS), Peripheral Nervous System (PNS), & Anatomical Structures in the CNS & PNS
Term
Central Nervous System
Definition
brain & spinal cord
Term
Peripheral Nervous System
Definition
neurons & axons of neurons outside the CNS (ganglia & peripheral nerves)
Term
anatomical structures in the CNS & PNS
Definition
nucleus, ganglion, tract, & nerves
Term
Nucleus
Definition
cluster of cell bodies of neurons involved in a similar function in the CNS
Term
Ganglion
Definition
cluster of neuron cell bodies in PNS involved in a similar function
Term
Tract
Definition
myelinated axons that run in parallel bundles in the CNS
Term
Nerves
Definition
myelinated axons that run in parallel bundles in the PNS
Term
Cranial Nerves
Definition
connected directly to the brain
Term
Spinal Nerves
Definition
run from various locations in the body to the spinal cord
Term
Sensory Transduction
Definition
incoming stimuli (chemical or physical stimuli from an animal's body or the external environment) are converted into neural signals
Term
Perception
Definition
conscious awareness of sensation; not all sensations are consciously perceived by an organism
Term
Sensory Receptor
Definition
recognizes stimulus & initiates signal transduction by creating graded potentials in the same or adjacent cells; either neurons or specialized epithelial cells; when the response is strong enough, an action potential is sent to the CNS
Term
Intensity of Sensory Stimuli
Definition
1. Amount of cell membrane depolarization directly related to intensity of stimulus (graded potential)
2. Membrane potential = Receptor potential
3. When a stimulus is strong enough, it will depolarize the membrane to the threshold potential & produce an action potential in a sensory neuron
4. Strength of the stimulus is indicated by the frequency of action potentials generated--strong stimulus generates more action potentials in a shorter amount of time
5. Brain interprets higher frequency of action potentials as a more intense stimulus
6. Different stimuli produce different sensations, respond because they activate specific neural pathways that are dedicated to processing only that type of stimulus
Term
Mechanoreceptors
Definition
Transduce mechanical energy
Term
Electromagnetic Receptors
Definition
Detect radiation within a wide range of the electromagnetic spectrum
Term
Photoreceptors
Definition
respond to visible light energy; electromagnetic receptor
Term
Thermoreceptors
Definition
respond to cold & heat; electromagnetic receptor
Term
Infared, Ultraviolet Receptors, & Magnetic Fields
Definition
electromagnetic receptors
Term
Chemoreceptors
Definition
respond to specific chemicals
Term
Nociceptors, or pain receptors
Definition
respond to extremes of heat, cold, & pressure, as well as to certain molecules such as acids; receptors with intense stimulus
Term
Mechanoreceptor
Definition
physically touching or deforming a mechanoreceptor opens ion channels in the plasma membrane: mechano-sensory proteins within the membrane
some mechanoreceptors are neurons & others are specialized epithelial cells
Term
Stretch receptors
Definition
example of mechanoreceptors
1. stretching alters proteins in cell membrane, causing ion channels, & causing membrane to depolarize
2. action potentials initiated, sent to the CNS
Term
Examples of stretch receptors
Definition
1. stretching of stomach wall is interpreted as fullness
2. stretching in blood vessels provides blood pressure info
Term
hair cells
Definition
example of mechanoreceptor
specialized epithelial cells
Term
deformable cilia resemble hairs
Definition
1. Ion channels open or close when cilia bend which changes membrane potential--frequency: influenced by the direction of bending
2. Membrane potential changes in hair cells, influence the fusion of neurotransmitter-containing vesicles
Neurotransmitters can trigger action potentials to
adjacent neurons
3. Found in the vertebrate ear & equilibrium organs, body surface of fish & some amphibians
Term
Skin receptors
Definition
mechanosensory system; Meissner's corpuscles & Pacinian corpuscles
Term
Meissner's corpuscles
Definition
sense touch & light pressure
lie just beneath skin surface
specialized dendritic branch
Term
Pacinian corpuscles
Definition
located much deeper beneath the surface
respond to deep pressure & vibration
specialized dendritic branch
Term
Lateral line system
Definition
mechanosensory systems; hair cells that detect changes in water currents; cilia of hair cells protrude into cupula structure within lateral line canal system; when cupula moves, cilia bend, & neurotransmitter released
Term
Audition (hearing)
Definition
mechanosensory system
Overview:
1. ability to detect & interpret sound waves
2. wavelength
3. frequency
4. short wavelengths have high frequencies perceived as high pitch or tone
5. long wavelengths have lower frequencies perceived as a lower pitch
Term
Wavelength
Definition
distance from the peak of one sound wave to the next
Term
Frequency
Definition
number of complete waves in a second (Hz)
Term
3 main compartments of Mammalian Ear
Definition
1. Outer ear
2. Middle ear
3. Inner ear
Term
Outer ear
Definition
pinna & auditory canal: separated from middle ear by eardrum
Term
Middle ear
Definition
ossicles (maileus, incus, stapes) connect eardrum to oval window
Term
Inner ear
Definition
cochlea (audition) & vestibular system (equilibrium, balance, proprioception)
Term
Eustachian tube
Definition
located in inner ear; connects to pharynx; equalizes pressure between middle ear & atmospheric pressure
Term
Movement of sound waves through the ear
Definition
1. sound waves enter the outer ear
2. tympanic membrane vibrates back & forth
3. ossicles transfer vibration to oval window
4. sends pressure waves through cochlea
5. waves travel from vestibular canal to tympanic canal & dissipate against round window
6. higher frequency sounds we hear pass through basilar membrane making it vibrate
Term
Transduction of sound waves in the Organ of Corti
Definition
1. Mechanical forces transduced into electrical signals by Organ of Corti
2. Hairs bending in one direction triggers neurotransmitter release
3. Action potentials triggered in dendrites of neurons of the auditory nerve
4. Frequency of action potentials determined by up-and-down vibration of basilar membrane
Term
Equilibrium or Proprioception
Definition
1. Ability to sense the position, orientation, & movement of the body
2. Many aquatic invertebrates have statocysts to send positional information: small, round chambers
Term
Vestibular system in vertebrates
Definition
1. Located in inner ear next to cochlea
2. Utricle & saccule detect linear movements of the head
3. Semicircular canals detect motion in 3D
Term
Utricle & saccule detect linear movements of the head
Definition
1. When head moves, inertia causes calcium-carbonate otoliths (which are embedded in gelatin substance to lag behind & bend cilia changing the membrane potential)
2. Utricle senses horizontal movements
3. Saccule senses vertical movements
Term
Semicircular canals detect motion in 3D
Definition
1. Hair cells embedded in gelatinous cupula
2. When head moves, fluid in the canal shift in the opposite direction pushing on cupula & bending hair cilia
3. Each canal oriented in one of 3 planes
Term
Electromagnetic sensing
Definition
detection of radiation within a wide range of the electromagnetic spectrum, including those wavelengths that correspond to visible light, UV light, & infared light, as well as electrical & magnetic stimuli
Term
Electromagnetic sensing examples
Definition
non-light systems:
1. Fish detect electrical signals from other fish
2. Platypus bill can detect electrical currents from prey
3. Homing pigeons use magnetite to accurately navigate
4. Pit vipers sense infared radiation (heat) from prey
Term
Photoreception
Definition
1. Photoreceptors detect photons of light arriving from the sun or other light source, or reflecting off an object
2. Photon-fundamental unit of electromagnetic radiation with the properties of both a particle & a wave
Term
Planaria photoreception
Definition
1. simple visual organ
2. eyecup containing endings of photoreceptor cells detects pressure or absence of light
3. layer of pigment casts shadows, so can detect direction of light
4. does not form visual images
Term
compound eyes of arthropods (& some annelids)
Definition
1. Many light detectors called ommatidia
2. Each light detection unit makes up one facet
3. Lens & crystalline cone focus light onto rhabdom
4. Retinula cells surrounding the rhabdom serve as photoreceptors
5. Pigment cells surround the rhabdom & retinula cells, keep light from one ommatidia from leaking to adjacent ommatidia
6. Extremely sensitive to movement, but less resolving power than single-lens eye
Term
Rhabdom
Definition
transparent tube
Term
Echinoderms
Definition
have eyespots on the tips of their arms; eyespots allow the organism to detect light & direction of light
Term
Single-lens eye
Definition
1. Found in vertebrates, some mollusks (squid & octopus), & in some snails & annelids
2. Light transmitted through pupil to retina at the back of the eye
3. Photoreceptors trigger electrical changes in neurons
4. Sclera
5. Cornea
6. Iris
Term
Sclera
Definition
strong outer connective tissue sheath
Term
Cornea
Definition
continuous with sclera but thin & clear
Term
Iris
Definition
pigmented smooth muscle controlling size of pupil
Term
Rods
Definition
1. Receptor cell in the human eye
2. sensitive to low intensity light
3. do not discriminate colors
4. used mostly at night
Term
structure of rods
Definition
1. a modified type of neuron
2. outer segment contains pigment
3. inner segment contains nucleus & other organelles
4. synaptic terminal
Term
synaptic terminal
Definition
neurotransmitter filled vesicles fuse with membrane
Term
Cones
Definition
1. Receptor cells in the human eye
2. require more light for stimulation
3. detect color
4. fewer cones than rods in human retina
Term
Complex of opsin protein in a complex with retinal
Definition
Retinal & several types of Opsin
Term
Retinal
Definition
vitamin A derivative that absorbs light energy
Term
Rhodopsin
Definition
type of opsin that is a rod pigment
Term
Cone pigments
Definition
type of opsin: humans have 3 (red, green, blue) distinct photopsins, some species have less, birds have 5
Term
Photons alter visual pigments
Definition
1. Photoreceptors differ from other sensory receptor cells because their membrane potential is in a slightly depolarized state (slightly positive membrane potential) when the cell is at rest
2. Depolarization in unstimulated state results in continuous release of glutamate
3. When exposed to light, retinal is altered
4. Opsin changes shape, intracellular signal is transmitted, resulting in membrane potential becoming hyperpolarization
5. Hyperpolarization is proportional to the intensity of light
Term
Chemoreception
Definition
1. Olfaction
2. Gustation
3. Chemicals bind to chemoreceptor cells, initiating signals that cause release of neurotransmitters from the chemoreceptor cell
4. Neurotransmitters initiate electrical responses in neurons that extend axons into the brain
Term
Olfaction
Definition
sense of smell
Term
Gustation
Definition
sense of taste
Term
Olfaction in mammals (chemoreception example)
Definition
1. Olfactory sensitivity on mammals is highly variable
2. Olfactory receptors have long cilia with specific receptors to bind odor molecules
3. Binding of chemical ligand results in action potentials being sent to olfactory bulb at the base of the brain
4. Overall smell sensation depends upon which set of olfactory receptor cells are activated.
Term
Olfactory sensitivity on mammals is highly variable
Definition
Depends on the density of olfactory receptor cells (5 million in humans v. 220 million in dogs)
Term
Olfactory receptors have long cilia with specific receptors to bind odor molecules
Definition
Only one type of odor receptor expressed per olfactory receptor cell
Term
Taste buds
Definition
1. clusters of chemosensory cells that detect particular molecules in food molecules dissolved in saliva
2. located in folds of tongue papillae
3. tips of sensory receptor cells in taste bud have microvilli that extend into taste pore
Term
Skeleton
Definition
structure or structures that serve one or more functions related to support, protection, & locomotion
Term
3 types of skeletons
Definition
1. hydrostatic
2. exoskeleton
3. endoskeleton
Term
Hydrostatic skeletons
Definition
1. water-filled cavity surrounded by muscle
2. water is nearly incompressible, so hydrostatic pressure can be used to extend parts of the body
Term
Hydrostatic skeletons in Cnidarians
Definition
body, tentacles can elongate or shorten
Term
Hydrostatic skeletons in Echinoderms
Definition
movement of tube feet
Term
Hydrostatic skeletons in Earthworms
Definition
move forward by passing a wave of muscular contractions along the length of the body: circular muscles squeeze & elongate while longitudinal
Term
Exoskeletons
Definition
1. external skeleton surrounding & protecting body
2. vary in complexity, thickness, & durability
3. Arthropods
4. Interior muscles connected to exoskeleton components for movement
Term
Exoskeletons in Arthropods
Definition
made of chitin, segmented for movement, must be shed to grow
Term
Endoskeletons
Definition
1. internal structures
2. do not protect body surface, only internal organs & other structures
Term
Examples of Endoskeletons
Definition
sponges (spicules), echinoderms (ossicles), & vertebrates
Term
2 parts of vertebrate skeleton
Definition
1. Axial
2. Appendicular
Term
Axial
Definition
main longitudinal axis (skull, spinal column, ribs)
Term
Appendicular
Definition
limb bones & girdles
Term
Joints of vertebrate skeleton
Definition
formed where 2 or more bones come together
Term
3 types of Joints
Definition
1. Pivot joints
2. Hinge joints
3. Ball-and-socket joint
Term
Pivot joints
Definition
neck bone--rotational movement
Term
Hinge joints
Definition
elbow & knee--movement in one plane
Term
Ball-and-socket joint
Definition
hip joint--movement in several planes
Term
Skeletal muscles
Definition
1. Muscle is a grouping of contractile cells (muscle fibers) bound together by connective tissue
2. Tendons link bones to skeletal muscle
3. Muscles, bones, & joints arranged in lever systems
4. Skeletal muscle tissue organization
Term
Muscles, bones, & joints arranged in lever system
Definition
1. lever system amplifies the velocity of muscle shortening
2. short, relatively slow movements of a muscle produces faster movements of the hand
Term
Skeletal muscle tissue organization
Definition
1. Skeletal muscle cells (muscle fibers) contain many parallel-arranged long protein fibers (myofibrils) along their length
2. Muscle cells are packaged in parallel into muscle bundles
3. A group of muscle bundles forms a muscle
Term
Each myofibril composed of a series of sarcomere unites
Definition
1. contractile
2. composed of interdigitated protein fibers
Term
Myofibrils
Definition
1. skeletal muscle is a type of striated muscle
2. striated muscle named for striped microscopic pattern of myofibrils
3. striated pattern reveals functional structure of contractile filaments and sarcomeres
4. sarcomeres are composed of interdigitating thick & thin filaments
Term
Myofibrils
Definition
1. skeletal muscle is a type of striated muscle
2. striated muscle named for striped microscopic pattern of myofibrils
3. striated pattern reveals functional structure of contractile filaments and sarcomeres
4. sarcomeres are composed of interdigitating thick & thin filaments
Term
Thick filaments
Definition
made of myosin protein--actively allow for contraction
Term
Thick filaments
Definition
made of myosin protein--actively allow for contraction
Term
Thin filaments
Definition
contain actin, troponin, & tropomyosin
Term
Thin filaments
Definition
contain actin, troponin, & tropomyosin
Term
Neuromuscular Junction
Definition
1. Junction of motor neuron's axon & muscle fiber
2. Axon branches into terminals
3. Region of muscle fiber under axon terminal is folded into junctional folds to increase surface area
4. ACh receptor is ligand-gated ion channel
5. Na+ flows into muscle cell leading to depolarization & an action potential
Term
Neuromuscular Junction
Definition
1. Junction of motor neuron's axon & muscle fiber
2. Axon branches into terminals
3. Region of muscle fiber under axon terminal is folded into junctional folds to increase surface area
4. ACh receptor is ligand-gated ion channel
5. Na+ flows into muscle cell leading to depolarization & an action potential
Term
Axon branches into terminals
Definition
terminals have stored vesicles containing neurotransmitter (acetylcholine)
Term
Axon branches into terminals
Definition
terminals have stored vesicles containing neurotransmitter (acetylcholine)
Term
Region of muscle fiber under axon terminal is folded into junctional folds to increase surface area
Definition
allows for there to be more acetylcholine receptors
Term
Region of muscle fiber under axon terminal is folded into junctional folds to increase surface area
Definition
allows for there to be more acetylcholine receptors
Term
Excitation-contraction coupling
Definition
1. Action potentials transmitted in muscle membrane trigger a rise in cytosilic Ca2+ released from sarcoplasmic reticulum
2. Triggers contraction
3. Ion pumps will return calcium to the sarcoplasmic reticulum, causing muscle to relax again
Term
Action potentials transmitted in muscle membrane trigger a rise in cytosilic Ca2+ released form sarcoplasmic reticulum
Definition
Transverse or T-tubules are invaginations of plasma membrane that conduct the action potential from the outer surface to inner regions
Term
Sarcoplasmic reticulum
Definition
endoplasmic reticulum is a specialized form
Term
Transport & Exchange
Definition
1. Some organisms are small enough that dissolved substances are able to diffuse in & out of their bodies--no transport system needed for efficient exchange
2. Primary function of circulatory systems is to transport necessary materials (O2, nutrients) to all the cells of an animal's body, & to transport waste products away from the cells where they can be released into the environment
Term
2 Basic types of transport/exchange systems
Definition
1. Gastrovascular cavities
2. Circulatory systems
Term
Gastrovascular cavities
Definition
1. Body cavity with a single opening to the outside
2. Cnidarians (jellyfish & hydras)
3. All of the animal's body cells are located near the cavity or slender extensions from it (tissue around outside)
4. Muscular efforts of the body wall cause movement of fluid increasing efficiency of exchange
Term
Circulatory systems
Definition
1. Transport fluid
2. Blood vessels
3. One or more pumps
Term
Open Circulatory System
Definition
1. Found in arthropods & some mollusks
2. Vessels connected to heart(s) (tubular) open into animal's body cavity
3. Fluid in vessels & interstitial fluid are the same
4. Nutrients & metabolic waste exchange by diffusion--between hemolymph--and body cells
5. In insects, no oxygen carrying pigments
6. Tracheal system is used for gas exchange
Term
Limitation for Open Circulatory System
Definition
hemolymph cannot be selectively directed to different tissues/areas of the body
Term
Closed Circulatory System
Definition
1. Blood & interstitial fluid are physically separated, only certain components exchanged between the two
2. Allows larger, more active animals higher to more efficiently pump blood to all body cells under high pressure
3. Found in annelids (earthworms), cephalopods (squids, octopus), & all vertebrates
Term
Common features of Closed Circulatory Systems
Definition
1. Blood (cells, large solutes) remains within vessels
2. One or more contractile, muscular hearts
3. Often blood proteins that bind to O2 & CO2, increasing carrying capacity
4. May contain disease-fighting cells & molecules
5. Flow can be adjusted to match local tissue/organ metabolic demands
6. Capacity to heal vessels when broken/wounded (clots)
Term
Blood
Definition
small solutes & water can move between vessels & Interstitial fluid
Term
Circulation found in animals
Definition
1. delivers nutrients & oxygen, picks up CO2 & waste products
2. deoxygenated blood is returned by veins to the heart
Term
Circulation found in annelids, fish, & cephalopods
Definition
single circulation
Term
Single circulation
Definition
1. Single blood circuit
2. Single atrium collects blood from tissues
3. SIngle ventricle pumps blood out of the heart
4. Arteries carry blood away from the heart to the gills
5. Blood picks up O2 & drops off CO2 & goes on through arteries to other body tissues (under low pressure)
Term
Circulation found in crocodiles, birds, & mammals
Definition
double circulation
Term
Double circulation
Definition
1. Two distinct blood circuits
2. Oxygenated & deoxygenated blood separates into 2 distinct circuits
3. Systematic circulation
4. Pulmonary circulation
5. 2 atria & 2 ventricles
Term
Systematic circulation
Definition
to the body
Term
Pulmonary circulation
Definition
to the lungs
Term
Major advantage of double circulation
Definition
high pressure circulation to 2 different systems
Term
Amphibians & reptiles
Definition
1. Rely on lungs & highly permeable skin to obtain O2 & get rid of CO2
2. Heart pumps blood to either pulmocutaneous circulation or systematic circulation
3. 2 atria to collect blood
4. Both atria dump into single ventricle
5. Internal structure causes oxygenated & deoxygenated blood to remain mostly separated
6. But, some mixing does occur reducing efficiency
7. Noncrodilian reptiles also have 2 atria & 1 ventricle but skin is not a major exchange surface
Term
Pulmocutaneous circulation
Definition
respiratory surfaces of lungs & skin
Term
Systematic circulation
Definition
body tissues within reptiles & amphibians
Term
Right atrium in reptiles & amphibians
Definition
blood that's been through the body (not lungs) & is low in O2 (except oxygenated blood from skin)
Term
Left atrium in reptiles & amphibians
Definition
blood from lungs (O2 rich when air breathing)
Term
Mammalian Heart
Definition
1. Septum separates atrai & ventricles
2. Blood enters from systematic or pulmonary veins into atrium
3. Through one-way atrioventricular (AV) valves into ventricles
4. Out one-way semilunar valves into systematic or pulmonary arteries
Term
Muscle contraction
Definition
1. Neurotransmitter release
2. Action potential conduction into T-tubules
3. Calcium release form sarcoplasmic reticulum
4. Calcium binding to troponin
5. Troponin effect tropomyosin, unblocking myosin binding sites on actin filaments
Term
Heart muscle is electrically excitable in Mammalian Heart
Definition
1. Mammalian heart can generate their own periodic action potentials--"myogenic" excitation
2. "Neurogenic" hearts of arthropods require regular electrical impulses from the nervous system
Term
"Myogenic" excitation
Definition
nervous input can increase or decrease rate
Term
Activation of Mammalian heart contraction
Definition
1. Action potential spreads because cardiac cells are electrically coupled by gap junctions (from a syncytium)
2. Both atria contract together forcing blood through AV valves into ventricles
3. Electrical impulses reach AV node conducts impulse to ventricles, after short delay
4. Both ventricles contract together forcing blood through semilunar valves into systematic or pulmonary arteries
Term
Both ventricles contract together
Definition
all valves shut so blood travels only in one way
Term
Sinoatrial node (SA)
Definition
pacemaker
Term
pacemaker
Definition
collection of modified cardiac cells that spontaneously & rhythmically generate action potentials
Term
Cardiac cycle
Definition
1. Events that produce a single heartbeat
2. Diastole & systole
3. Heart valves open & shut in response to pressure gradients
Term
Diastole
Definition
atria contract & ventricles fill (systematic blood pressure lowest)
Term
Systole
Definition
ventricles contract & blood is ejected from the heart (systematic blood pressure highest)
Term
Blood
Definition
fluid connective tissue in closed circulatory system: transport medium
Term
4 prominent components of vertebrate blood
Definition
1. Plasma
2. Erythrocytes
3. Platelets or thrombocytes
4. Leukocytes
Term
Plasma
Definition
water & solutes
1. function in buffering, water balance, & cell transport
2. contains dissolved proteins, gases, minerals, nutrients
Term
Erythrocytes
Definition
red blood cells
Term
Red Blood Cells (RBCs)
Definition
oxygen transport using hemoglobin
Term
Platelets & thrombocytes
Definition
role in formation of blood clots (fibrin precipitation)
Term
Leukocytes
Definition
white blood cells
Term
White Blood Cells (WBCs)
Definition
defend body against infection & disease
Term
Arteries
Definition
1. conduct blood away from the heart
2. layers of smooth muscle & elastic connective tissue around smooth endothelium (epithelial tissue on inside)
Term
Arterioles
Definition
1. smaller in diameter, branches of arteries
2. walls thinner than arteries, lack thick layer of connective tissue
3. have smooth muscle encircling
Term
Smooth muscle encircling arterioles
Definition
1. Can dilate or constrict to control blood distribution to tissues
2. A key factor in blood pressure regulation
Term
Capillaries
Definition
1. Smallest & narrowest, thinnest walled vessels in the body
2. Arterioles branch into fine capillary networks
3. Site of gas & nutrient/waste exchange
4. Single-celled thick endothelium layer on a basement membrane
Term
Arterioles branch into fine capillary networks
Definition
most cells in the body are within a few cell diameters of a capillary
Term
Single-celled thick endothelium layer on a basement membrane
Definition
1. Fenestrated capillaries have opening or fenestrations, allow for movement of considerable water & small solutes through the walls
2. Continuous capillaries have smooth walls with no fenestrations; permit less water & solute movement
Term
Movement of materials through capillaries
Definition
1. Blood enters capillary on arteriole end under hydrostatic pressure
2. Pressure forces some fluid out of the blood (not RBCs or large proteins)
3. Hydrostatic pressure decreases along the capillary bed
4. Proteins in the blood create an osmotic force that draws fluid back into blood
5. Most of the fluid that leaves will be recaptured by the venule end of the capillary
6. Lymphatic system will collect fluid that is not captured & return it to the blood
Term
3 different ways pressure forces some fluid out of the blood
Definition
1. between cells
2. small pores
3. fenestrations
Term
Venules
Definition
1. capillaries come together to drain into venules
2. thin walls
Term
Veins
Definition
1. Thinner & less elastic than arteries
2. Need help returning blood to the heart
Term
3 ways veins need help returning blood to the heart
Definition
1. smooth muscle contractions help propel blood
2. veins squeeze by skeletal muscles
3. flow is directed by unidirectional valves: changes in pressure
Term
Adaptative capabilities of the vascular system
Definition
1. System must adapt to changing conditions
2. Blood can be routed to different areas in proportion to their need for O2 & nutrients
Term
Vascular system must adapt to changing conditions such as:
Definition
sleep, feeding, sudden activity, & emergencies
Term
Blood can be routed to different areas in proportion to their need for O2 & nutrients
Definition
1. Can be controlled by vasodilation or vasoconstruction
2. "Precapillary sphincters"
Term
Respiration, Gas Exchange
Definition
refers to gas exchange processes in the body
Term
2 types of Respiration, Gas Exchange
Definition
1. Pulmonary respiration
2. Internal Respiration
Term
Pulmonary Respiration
Definition
gas exchange moves carbon dioxide & oxygen between the air & blood
Term
Internal Respiration
Definition
gas exchange moves carbon dioxide & oxygen between blood & cells
Term
Air composition
Definition
21% oxygen
78% nitrogen
>1% CO2 & other gases
Term
Atmospheric Pressure
Definition
1. pressure exerted by the atmosphere on the body surfaces of animals
2. decreases at higher elevations
Term
Atmospheric Gases
Definition
Measure in mmHg or kPa: -1kPa = 7.5 mmHg
Term
Sea Level Atmospheric Gases
Definition
760 mmHg
Term
Partial Pressures
Definition
1. Atmospheric pressure is the sum of the "partial pressure" exerted by each gas in air
2. Partial pressure is determined by the proportion of a particular gas in the atmosphere
3. Direction of gas diffusion is driven by partial pressure gradients
4. Partial pressure can be defined for dissolved gases (gases dissolved in water, blood, hemolymph, etc.)
5. Rate & direction of gas diffusion into or out of blood is determined by partial pressure differences
Term
Proportion of a particular gas in the atmosphere
Definition
PO2 = 0.21 x 760mmHg = 160mmHg
Term
Solubility of gases in water
Definition
1. Gases dissolve in solution-freshwater, seawater, or body fluids
2. Most gases dissolve poorly in water
Term
Factors influencing solubility in water
Definition
1. Higher pressures will result in more gas in solution up to a limit for each gas
2. Cold water holds more gas than warm water
3. The pressure of other solutes decreases the amounts of gas that dissolves into solution
Term
All respiratory organs share common features for gas exchange
Definition
1. Moist surfaces in which gases dissolve & diffuse
2. Often structured to provide large surface area for gas exchange
3. Extensive blood supply
Term
Challenges for terrestrial vs. aquatic animals
Definition
1. Aquatic animals have less available oxygen (partial pressure of O2 less in water than in air)
2. When temperature changes in water, O2 availability also fluctuates (gases dissolve more easily in cold water than in warm water)
3. Terrestrial animals have to deal with desiccation (drying out of respiratory membranes)
4. Water is denser than air
Term
Invertebrates with one or a few cell layers can use diffusion for gas exchange
Definition
1. Some do not even need specialized transport mechanism
2. Small, flat body form
Term
Larger organisms
Definition
1. Body surfaces may be permeable to gases
2. Specialized exchange surfaces
Term
Body surfaces may be permeable to gases in larger organisms
Definition
amphibians are the only vertebrate to rely on their skin for gas exchange under water
Term
Specialized exchange surfaces within larger organisms
Definition
gills, tracheae (insects), lungs
Term
External gills
Definition
1. Vary widely in appearance but all have a large surface area (extensive projections)
2. May exist in one body area or be scattered over a large area, branching
Term
Limitations of external gills
Definition
1. Unprotected & subjected to damage
2. Appearance & motion may attract predators
Term
Internal gills
Definition
1. Fish gills are confined & protected within opercular cavity covered by operculum
2. Gill arches
3. Filaments branch off of gill arches
4. Lameilae branch off of filaments
5. Blood vessels run the length of the filaments
Term
Gill arches
Definition
main support structure in internal gills
Term
Blood vessels run the length of the filaments in internal gills
Definition
1. oxygen-poor blood travels through afferent vessel
2. oxygen-rich blood travels through efferent vessel
Term
Blood in capillaries flows in opposite direction as water flow-allows for more efficient gas exchange
Definition
counter-current exchange
Term
Counter-current exchange
Definition
1. An intermediate equilibrium state is reached [concurrent]
2. Disequilibrium maintained; exchange continues over; entire exchange surface [countercurrent]
Term
Concurrent
Definition
concentration is off
Term
Countercurrent
Definition
Concentration is more equal
Term
Insect trachea
Definition
1. Spiracles on the body surface lead to trachea that branch into tracheoles terminating near every body cell
2. Small amount of fluid for gas to diffuse into
3. Muscular movements of body draw air into & out of trachea
4. Open circulatory system of insect not used in gas exchange
5. Oxygen diffuses directly from air into tracheae to tracheoles to body cells
6. Very efficient-supports insect flight muscle with highest metabolic rate known
Term
Lungs
Definition
1. Almost all air-breathing terrestrial vertebrates use lungs
2. Arachnids (scorpions & some spiders) have book lungs that actually more closely resemble gills
3. Lungs may be filled using positive or negative pressure
4. Lungs can be ventilated using tidal or flow-through systems
Term
Positive pressure in lungs
Definition
pushing air
Term
Negative pressure in lungs
Definition
pulling air
Term
Tidal systems in lungs
Definition
pulled in & pushed out
Term
Flow-through systems in lungs
Definition
doesn't follow same path inward & outward
Term
Amphibian Lungs & Positive Pressure Filling/Breathing
Definition
1. Most amphibians have lungs that are simple sacs
2. Relatively low surface area
3. Ventilate lungs similar to buccal pumping of fish
4. Lowers bottom jaw to create pressure gradient to suck air in
5. Muscles in mouth constrict (with mouth closed) to raise pressure & force air into lungs-positive pressure filling
6. A few species of reptiles also use positive pressure filling
Term
Negative Pressure Ventilation
Definition
1. Reptiles, birds, mammals
2. Volume of thoracic cavity expands, creating negative pressure, & air drawn into lungs
Term
Mammals = tidal ventilation
Definition
1. Inhalation
2. Exhalation
Term
Inhalation
Definition
1. intercostals contract to move chest wall up & out
2. diaphragm contracts & drops down-thoracic cavity enlarges
3. pressure decreases, air sucked in
Term
Exhalation
Definition
1. intercostals & diaphragm relax-thoracic cavity compressed
2. pressure increases, air pushe out
Term
Avian lungs & flow-thru breathing
Definition
1. During inhalation, air moves through trachea into posterior air sacs, not lungs (base of body)
2. During exhalation, air moves through air sacs (pariabronchi) into lungs, then pushed through trachea
3. Air sacs expand & shrink--do not participate in gas exchange
4. Blood flows crosscurrent with respect to air movement
Term
During inhalation in avian lungs
Definition
air in lungs (from previous breath) moves into anterior air sacs
Term
During exhalation in avian lungs
Definition
1. Region of gas exchange, single direction flow through lung into anterior air sacs; exhalation empties air from these sacs
2. Air in anterior sacs exits body
Term
Mammalian Respiratory System Anatomy
Definition
1. Nose & mouth
2. Mucus in the nose cleans the air of dust
3. Pharynx
4. Larynx
5. Trachea
6. Lungs
7. Trachea branches into 2 bronchi
8. Bronchioles
9. Alveoli
Term
Nose & mouth
Definition
air is warmed & humidified
Term
Pharynx
Definition
the hollow tube about 5 in. long that starts behind the nose & ends at the top of the trachea (windpipe) & esophagus (tube that goes to stomach)
Term
Larynx
Definition
vocal cords
Term
Trachea
Definition
glottis (opening to trachea) protected by epiglottis, rings of cartilage (prevent collapse during inhalation), cilia & mucus trap particles
Term
Bronchioles
Definition
surrounded by circular muscle to dilate or constrict passage
Term
Alveoli
Definition
1. small pouches
2. site of gas exchange
3. one cell thick walls
4. surfactant chemicals reduce surface tension, prevent alveoli from collapsing
5. inner surface coated with extracellular fluid for gases to dissolve
Term
Gas exchanges in body
Definition
differences in pressures are going to drive diffusion of the gases in the proper directions
Term
water
Definition
1. a major portion of the body mass
2. solvent for chemical reactions
3. transport vehicle
Term
Dehydration may comprise essential processes
Definition
E.g. the circulatory system & regulation for body temperature
Term
circulatory system & regulation for body temperature
Definition
1. sweat draws off heat energy
2. taking water to surface of skin-taking heat
Term
Electrolytes
Definition
1. salts dissociate in solution into charged ions
2. electrolyte balance important
3. imbalance can alter membrane potential or disrupt other cellular activities
Term
Osmosis
Definition
1. water moves between adjacent body compartments by osmosis down an osmotic gradient
2. changes in salt concentration in one compartment will lead to change in fluid distribution between compartments
3. shrinking or swelling cells can rupture plasma membrane leading to cell death
4. change of osmosis changes water content of cells
Term
water
Definition
1. a major portion of the body mass
2. solvent for chemical reactions
3. transport vehicle
Term
Dehydration may comprise essential processes
Definition
E.g. the circulatory system & regulation of body temperature
Term
Circulatory system & regulation of body temperature
Definition
1. sweat draws off heat energy
2. taking water to surface of skin-taking heat
Term
Electrolytes
Definition
1. Salts dissociate in solution into charged ions
2. Electrolyte balance important
3. Imbalance can alter membrane potential or disrupt other cellular activities
Term
Osmosis
Definition
1. Water moves between adjacent body compartments by osmosis down an osmotic gradient
2. Changes in salt concentration in one compartment will lead to change in fluid distribution between compartment
3. Shrinking or swelling cells can rupture plasma membrane leading to cell death
4. Change of osmosis changes water content of cells
Term
Obligatory Exchanges
Definition
many vital processes have the potential to alter salt & water balance
Term
Breathing
Definition
1. loss of water by evaporation
2. fish can lose or gain water as water passes over gills
Term
Cellular Respiration
Definition
production of metabolic water: glucose + oxygen = carbon dioxide + water
Term
Temperature Regulation
Definition
panting & sweating cause loss of water & salts
Term
Elimination of wastes
Definition
salts & water are lost in feces & urine
Term
Food intake
Definition
salt & water content
Term
Osmolarity of body fluids compared to fresh & saltwater environments
Definition
1. Osmolarity
2. Internal fluid osmolarity of most fish & other vertebrates around 225-400 mosm/L
3. Freshwater less than 25 mosm/L
4. Seawater 1000 mosm/L
Term
Osmolarity
Definition
Total concentration of dissolved solute molecules/Liter:
150 mM NaCl solution = 300 mosm/L
one Na+ & one Cl- made when salt dissociates
Term
Water Balance in Aquatic Organisms
Definition
water moves by osmosis from areas of low osmolarity to high osmolarity
Term
Osmoregulation in Freshwater Fish
Definition
1. Gain water when ventilating gills
2. Kidneys reproduce copious dilute urine
3. Specialized gill epithelial cells transport Na+ & Cl- from water into fish's capillaries
Term
Kidneys in freshwater fish reproduce copious dilute urine
Definition
don't drink much because comes on by osmosis
Term
Osmoregulation in Saltwater Fish
Definition
1. Lose water across gills
2. Produce very little urine
3. Drink seawater to replace water lost
4. Expend energy to transport excess salt out of body through gill epithelial cells
Term
Osmoconformers
Definition
many marine invertebrates & cartilaginous fishes (sharks, rays) maintain body fluid osmolarity similar to the surrounding seawater
Term
Invertebrates
Definition
many are osmoconformers, with wide tolerance (i.e. tidal pool organisms)
1. often have similar ionic concentration as seawater
2. hagfish are the only vertebrate that exhibit this approach to water balance
Term
Sharks
Definition
allow urea to accumulate in blood
1. Total solute concentration similar to seawater, but solute composition different
Term
Nitrogenous wastes
Definition
1. Product of protein & nucleic acid degradation
2. Toxic at high concentrations
3. Must be eliminated from the body
Term
Nitrogenous wastes are eliminated from the body
Definition
1. various organs & organ systems utilized
2. the chemical forms of excretal nitrogenous wastes among animals varies
Term
Ammonia (NH3) & ammonium ions (NH4+)
Definition
1. Most toxic of nitrogenous wastes
2. Many aquatic animals can excrete it as soon as it forms
3. Some terrestrial snails & crustaceans secrete it as a gas
4. Chief advantage is that energy is not required for production
Term
Urea
Definition
1. All mammals, some marine fishes, some reptiles, & some terrestrial invertebrates
2. Less toxic, so doesn't need as large a volume of water for excretion
3. Can tolerate some urea accumulation
4. Drawback is conversion of ammonia to urea requires ATP
Term
Uric Acid
Definition
1. Birds, insects, & most reptiles
2. Less toxic than ammonia
3. More energetically costly than urea to make from ammonia
4. Balanced against water conserved by excreting semisolid, partly dried precipitate
Term
Vertebrate Urinary System
Definition
1. Organ system designed to remove and dispose of nitrogenous wastes, excess water, & blood solutes
2. For terrestrial organisms, water conservation is often important. The vertebrate urinary system provides a mechanism to concentrate waste products before release to reduce water loss.
Term
Organs, Important Structures in Vertebrate Urinary System
Definition
1. Kidney
2. Ureters
3. Urinary Bladder
4. Urethra
Term
3 Main Structures of Kidney
Definition
1. Renal Cortex
2. Renal Medulla
3. Renal Pelvis
Term
Excretory Organs Function
Definition
1. Filtration
2. Reabsorption
3. Secretion
4. Excretion
Term
Filtration
Definition
1. organ acts like a filter to remove water & small solutes from blood while leaving behind blood cells & large solutes
2. produces filtrate that is moved into excretory tubules
Term
Reabsorption
Definition
desirable material in filtrate is recaptured & retrained to blood
Term
Secretion
Definition
additional solutes moved into filtrate by active transport mechanisms
Term
Excretion
Definition
filtrate containing waste products & non-reabsorbed solutes released from the body
Term
Photonephridia
Definition
1. found in Platyhelminthes
2. Simplest filtration mechanism in invetebrates
3. Primarily osmoregulatory
Term
Simplest filtration mechanism in invertebrates
Definition
1. Series of branching tubules filters fluids from body cavity using beating of ciliated cells (flame cells)
2. Excess water & some wastes emptied through openings in body walled called nephridiopores
3. Beneficial solutes reabsorbed prior to excretion
Term
Primarily Osmoregulatory
Definition
1. Help to rid the body of excess water
2. In these organisms, nitrogenous wastes mainly diffuse out of the body
Term
Metanephridia
Definition
1. Found in Annelids
2. Pairs located in each body segment
3. Tubular network beginning in funnel-like structure called nephrostome
4. Collect coelomic fluid containing nitrogenous wastes, dissolved solutes
5. Beneficial solutes reabsorbed along the tubule length, unneeded solutes remain in filtrate
6. Nitrogenous wastes, excess solutes excreted through nephridiopores in body wall
Term
Malpighian tubules
Definition
1. Found in insects
2. Not a filtration system
3. Tubules extend from the intestine
4. Cells lining tubules actively transport & secrete solutes & uric acid from hemolymph into lumen of the malpighian tubules
5. Transport of solutes creates osmotic gradient, drawing water into tubule
6. Material in tubules move into hindgut where water and beneficial solutes are reabsorbed
7. Nitrogenous wastes & others excreted together with feces through anus
Term
Nephrons in Vertebrate Kidney Function
Definition
1. Specialized tubules composed of epithelial cells that actively transport sodium and other ions for salt and water homeostasis and nitrogenous waste elimination
2. Nephrons are the functional of the kidney
3. As many as several million in each kidney
4. A nephron consists of renal corpuscle & tubule
5. Nephron tubules empty into collecting duct
6. Glomerulus
7. Bowman's capsule encloses Bowman's space
8. Filtration
Term
Renal corpuscle
Definition
forms area of filtration
Term
Tubule
Definition
performs secretion and reabsorption
Term
Glomerulus
Definition
1. Cluster of interconnected fenestrated capillaries
2. Supplied by afferent arteriole
3. Drained by efferent arteriole
4. Podocytes form filtration slits (physical barriers)
Term
Bowman's space
Definition
space in between Bowman's capsule & glomerulus
Term
Filtration in Nephrons
Definition
1. As blood flows through glomerulus, about 20% of plasma leaves capillaries and filters into Bowman's space
2. Proteins & blood cells remain in plasma
3. Glomerular filtrate = water & small solutes
4. Glomerular filtration rate (GFR) is rate of filtrate production by kidneys
Term
Nephron Tubule
Definition
1. Continuous with Bowman's capsule
2. Epithelial cells differ in structure and function along length
3. Segments of loop of Henle
4. Distal convoluted tubules empty into collecting ducts
5. Tubule surrounded by peritubular capillaries near junction of cortex & medulla and vasa recta capillaries in the medulla
6. Processes occurring in nephron tubules
Term
Segments of Loop of Henle
Definition
1. Proximal convoluted tubule--proximal to Bowman's capsule
2. Loop of Henle
3. Distal Convoluted Tubule
Term
Loop of Henle
Definition
1. descending goes down into medulla
2. ascending comes up out of the medulla
Term
Processes occurring in nephron tubules
Definition
1. Reabsorption of proximal tubule
2. Water & salt movement in the Loop of Henle
Term
Reabsorption of proximal tubule
Definition
1. Filtrate is around 300 mosm/L similar to blood
2. Water & solutes are reabsorbed into peritubular capillaries to return to body
3. Depending on solute, 2/3 to all is reabsorbed
Term
Various solutes that are reabsorbed in nephron tubule
Definition
1. Na+
2. Cl-
3. K+
4. HCO3-
5. organic molecules like glucose & amino acids
Term
Water and salt movement in the Loop of Henle
Definition
1. Descending loop permeable to water but not solutes
2. Ascending loop (thick segment) not permeable to water & actively transports salts out
Term
Descending loop in Loop of Henle permeable to water but not solutes
Definition
1. Water leaves by osmosis because surrounding fluid hyperosmotic
2. Fluid increases in osmolarity & solute concentration
Term
Ascending loop (thick segment) in Loop of Henle not permeable to water & actively transports salts out
Definition
1. Tubule fluid decreases in osmolarity
2. Interstitial fluid increases in osmolarity (contributing to hyperosmotic fluid around descending loop)
Term
Concentration of Urine/Conservation of Water
Definition
1. An Osmotic Gradient in the Kidney medulla is key to concentrating urine and conserving water
2. Establishing the osmotic gradient in the kidney medulla
Term
An Osmotic Gradient in the Kidney medulla is key to concentrating urine & conserving water
Definition
1. Osmotic gradient, lower solute concentration in cortex and outer medulla, higher solute concentration in inner medulla
2. Collecting duct permeable to water but not to most ions
3. Osmotic movement of water from the collecting ducts to interstitial fluid of the medulla
Term
Establishing the osmotic gradient of the kidney medulla causes:
Definition
1. Pumping of NaCl out of ascending limb of loop of Henle
2. Facilitated diffusion of urea out of lower collecting duct
Term
Pumping of NaCl out of ascending limb of loop of Henle
Definition
causes osmosis of water out of descending loop & collecting ducts
Term
Facilitated diffusion of urea out of lower collecting duct
Definition
concentration of urine due to water loss by osmosis increases collecting duct urea concentration
Term
Ecology
Definition
The study of the interactions between organisms and between organisms and their environments
Term
Levels of Ecological Study
Definition
1. Organismal
2. Population
3. Community
4. Ecosystems
Term
Organisms Ecology
Definition
1. Focuses on the specific characteristics of individual organisms, and how these characteristics contribute to survival
2. Physiological adaptations
3. Behavioral patterns
4. Mating
Term
Physiological adaptations within organismal ecology
Definition
tolerance/adaptation to changes in environmental conditions (temperature, water salinity)
Term
Behavioral patterns within organismal ecology
Definition
1. Migration
2. Foraging behaviors-how look for food
3. Social interactions
Term
Social Interactions within organismal ecology
Definition
1. Territoriality
2. Communication between individuals
3. Living in groups (herds, hives, flocks, etc.)
4. Altruism
5. Mating
Term
Mating within organismal ecology
Definition
1. Seasonal patterns
2. Mate choice
3. Mating behaviors, structures
Term
Population Ecology
Definition
Focuses on the factors that influence the size, density, or growth of populations
Term
Population
Definition
group of interbreeding individuals (same species) present in a particular area
Term
Community Ecology
Definition
focuses on how groups of different species interact, and form communities with distinct characteristics; can be descriptive
Term
Community
Definition
a group of populations of different species that live in the same place at the same time
Term
Ecosystems Ecology
Definition
1. Focuses on the movement of energy and materials through organisms and their communities
2. Nutrient cycles
Term
Ecosystem
Definition
All the organisms in an area (biotic component) along with the abiotic environment (soil, water, nutrients, gases, ...)
Term
Food chains & webs
Definition
1. producers & consumers
2. trophic level transfer efficiencies
Term
Trophic Level Transfer Efficiencies
Definition
What proportion of energy from one level in a food web is assimilated in the next higher level?
Term
Nutrient Cycles
Definition
1. Carbon cycles
2. Nitrogen cycles
3. Sulfur cycles
4. Water cycles
Term
General Aspects of Population Ecology
Definition
1. Uses tools of demography
2. Density
3. Population growth affects population density
Term
Tools of demography
Definition
1. birth rates
2. death rates
3. age distributions
4. sizes of populations
Term
Density
Definition
number of organisms in a given unit area
Term
Quantifying Population Density
Definition
1. Simple visual count
2. Sampling methods to extrapolate captured organism
3. Mark-recapture method
Term
Mark-recapture method
Definition
# of marked ind. in 1st chain/total pop. size, N = # of marked recaptures in 2nd catch/total # of 2nd catch

total pop. size, N = # of ind. 1st catch x Total # of 2nd catch/# of marked recaptures in 2nd catch
Term
Dispersion Patterns
Definition
1. Clumped
2. Uniform
3. Random
Term
Clumped
Definition
1. most common
2. resources tend to be clustered in nature
3. social behavior may also promote the pattern
Term
Uniform
Definition
1. competition may cause this pattern
2. may also result from social interactions
Term
Random
Definition
1. rarest
2. resources are rarely randomly spaced
3. may occur where resources where resources are common & abundant
Term
Survivorship Curve definition
Definition
plots numbers of surviving individuals (or proportion) for a particular age cohort
Term
Survivorship Curve
Definition
May use logs to make it easier to examine wide range of population sizes
Term
3 common patterns of survivorship curves
Definition
1. Type 1
2. Type 2
3. Type 3
Term
Type 1
Definition
rate of loss of juveniles low & most individuals survive later in life
Term
Type 2
Definition
fairly uniform death rate over the lifespan
Term
Type 3
Definition
rate of loss for juveniles high & then low for survivors
Term
Life tables
Definition
1. Data collected on the number of individuals alive in each defined age class
2. Males often not included (usually not a limiting factor for population growth)
Term
North American beaver example
Definition
1. Trappers provided mandibles
2. Teeth analyzed for age classification
3. Assembled a "state" life table
Term
Age-specific fertility rate, Mx
Definition
1. proportion of female offspring born to females of particular age
2. 100 females of a certain age produce 75 female offspring Mx=0.75
Term
Age-specific survivors hip rate, Ix
Definition
1. proportion of individuals that will live to the next year
2. IxMx = contribution of each age class to the overall population growth
Term
Estimating Reproductive Rates from Life Table Data
Definition
1. Rsub0 = net reproductive rate
2. To calculate future size of population in the next generation, multiply # of individuals in the population by the net reproductive rate
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