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Exam 2
Unit 2 of Ecol 182 with Robichaux; only need to study up to 10 Molecular. REQUIRED READING: 725-735, 795-802 (ROOTS****)
80
Biology
Undergraduate 1
02/23/2012

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Term
Plants are multicellular, _____________ organisms that have __________ with cellulose.
Definition
photosynthetic; cell walls
Term
Plants retain their ____________ within the tissues of the maternal parent where they develop into _____________.
Definition
zygotes; embryos
Term
What is the key evolutionary innovation that enables plants to colonize land?
Definition
The developing embryo of a plant is protected from desiccation.
Term
Plants form a ______________ lineage (i.e., they are all descendants of a common ancestor that made the transition from an aquatic to a terrestrial environment).
Definition
monophyletic
Term
DNA sequence data indicate that the ancestors of plants belong to the _____________ lineage, which is a lineage of green algae.
Definition
charophyte
Term
Two key processes in the sexual life cycles are __________ (or reduction division) and ______________.
Definition
meiosis; fertilization
Term
Sexual reproduction generates ______________, which is important in evolution.
Definition
genetic variation
Term
In the sexual life cycles of humans and many other animals, the gametes are __________ products of ___________. In other words, in the haploid phase of the life cycle, there is not an intervening _______________ stage in humans, as there are in plants.
Definition
direct; meiosis; multicellular
Term
During the diversification of plants, there has been a shift from life cycles in which the ___________ is larger, longer-lived, and more self-sufficient to those in which the ___________ is larger, longer-lived, and more self-sufficient.

In the comparisons which plants is this shift the most evident?
Definition
gametophyte; sporophyte; mosses, ferns, and gymnosperms
Term
In the typical _________ life cycle, the multicellular gametophyte is photosynthetic and free-living.
Definition
moss
Term
In the typical ________ life cycle, both the multicellular gametophyte and multicellular sporophyte are photosynthetic and free-living
Definition
fern
Term
In the typical _____________ life cycle, the multicellular sporophyte is photosynthetic and free-living
Definition
gymnosperm
Term
In the gymnosperm life cycle, two distinct types of spores are produced by meiosis, called _____________ and _____________.
Definition
microspores; megaspores
Term
____________ give rise via mitosis to multicellular male gametophytes, which then give rise to the male gametes (i.e., sperm).
Definition
microspores
Term
______________ give rise via mitosis to multicellular female gametophytes, which then give rise to the female gametes (i.e., eggs).
Definition
megaspores
Term
There are more than ___________ known species of angiosperms, which is by far the largest number for any plant group.
Definition
230,000
Term
In plants, the haploid phase of the life cycle is called __________ generation and, the diploid phase is called the __________ generation. Thus the life cycles of plants are said to involve "______________".
Definition
gametophyte; sporophyte; alternation of generations
Term
Gymnosperms are thus said to be ____________.
By contrast, mosses are ___________, as are most (but not all) ferns.
Definition
heterosporous; homosporous
Term
In terms of their ecological and economic importance, angiosperms are _____________. ________________. and _____________.
Definition
primary producers; sources of food; sources of drugs
Term
Why are angiosperms primary producers?
Definition
In most terrestrial ecosystems, angiosperms account for most of the photosynthetic conversion of radiant energy into chemical energy. Angiosperms thus form the basis for most terrestrial food chains.
Term
What are some examples of food that uses angiosperms?
Definition
-all cereal grains and most vegetables, fruits, and nuts are obtained from angiosperms,

- most spices are obtained from angiosperms, and

- most non-alcoholic and alcoholic beverages are made from angiosperms.
Term
How are angiosperms sources of drugs?
Definition
Many angiosperms produce chemicals with significant medicinal properties. Other angiosperms produce chemicals with significant effects on the human central nervous system.
Term
What are examples of chemicals made with angiosperms and what are their medical properties?
Definition
Quinine – used in treating malaria

Vincristine – used in treating leukemia

Curare – used as muscle relaxant in open-heart surgery

Diosgenin – used as a precursor to the active ingredient in oral contraceptives
Term
What are chemicals made with angiosperms that affect the human central nervous system?
Definition
Mescaline – from peyote cacti (Lophophora)

Opium – from poppies (Papaver)

Cocaine – from cocoa plants (Erythroxylum)

Caffeine – from coffee plants (Coffea)
Term
In angiosperms, the ovules are enclosed within specialized structures called _________ (or pistils), which are contained in the _________.
Definition
carpels; flowers
Term
Angiosperms exhibit __________ fertilization and _________ formation.
Definition
double; endosperm
Term
Since angiosperms are heterosporous, the multicellular male gametophyte is the ___________, and the multicellular female gametophyte is the ____________.
Definition
pollen grain; embryo sac
Term
The vegetative body of a typical angiosperm consists of __________, ___________, and ___________.
Definition
roots; stems; leaves
Term
___________ are involved in water and nutrient uptake and in support (or anchoring).
Definition
roots
Term
_________ are involved in the transport of water, nutrients, sugars, etc. between the roots and leaves and in support.
Definition
stems
Term
__________ carry out photosynthesis.
Definition
leaves
Term
_________ is involved in the transport of water and nutrients, and ____________ is involved in the transport of sugars produced by photosynthesis of other compounds.
Definition
xylem; phloem
Term
The upper and lower leaf surfaces are covered by a ______________ to prevent water vapor from diffusing out of the leaf.
Definition
waxy cuticle
Term
The waxy cuticle prevents ________ from diffusing into the leaf. This is a major dilemma because the plant needs this for photosynthesis.
Definition
CO2
Term
The _________ (or pore, located in the leaf epidermis) is surrounded by two specialized epidermal cells called __________.
Definition
stoma; guard cells
Term
What are two physical features of the guard cells that are important to the mechanism of stomatal opening?
Definition
1. Specialized bands of inelastic cellulose microfibrils are oriented perpendicular to the long axis of each guard cell, and

2. The cell wall of each guard is thicker on the side close to the stoma than on the side away from the stoma.
Term
What does the basic mechanism of stomatal opening (in sequence) involve?
Definition
1. active transport causing potassium ions to move into the guard cells from the surrounding epidermal cells (remember: active transport requires energy),

2. net movement of water into the guard cells to maintain osmotic balance,

3. increased turgidity of the guard cells, and

4. stomatal opening.
Term
A reverse process, first involving ___________ of potassium ions out of the guard cells into the surrounding epidermal cells, results in stomatal ________.
Definition
passive diffusion; closing
Term
Stomata regulates the balance between CO2 uptake and _________ loss.
Definition
water vapor
Term
A key process that determines how plants respond in different environments is ______________.
Definition
photosynthesis
Term
Plants obtain all of their chemical energy from _____________ once their seed resources have been depleted.
Definition
photosynthesis
Term
Photosynthesis may be divided into two major sets of reactions that occur in the _____________.
Definition
chloroplasts
Term
The _____________ (also known simply as the “light reactions”) occur in the thylakoid membranes in the chloroplasts.
Definition
light-harvesting reactions
Term
Light energy in the form of photons is absorbed by ____________ and associated accessory pigments.
Definition
chlorophyll
Term
The light energy is converted into chemical energy in the form of _____ and _______.
Definition
ATP; NADPH
Term
____________ conversion occurs via a complex series of physical and chemical reactions involving an initial electron donor (H2O), two photosystems (I and II), an electron transport chain, and a final electron acceptor (NADP+).
Definition
Light energy
Term
The ______________ (also known as the “dark reactions”) occur in the stroma (or protein-rich matrix) in the chloroplasts.
Definition
CO2-fixation reactions
Term
The ________________ consist of a primary reaction by which CO2 is chemically bound (or “fixed”), together with a series of chemical reactions that ultimately yield glucose as a product and that serve to regenerate the primary CO2-acceptor molecule.
Definition
CO2-fixation reactions
Term
This series of reactions uses the ATP and NADPH generated by the light-harvesting reactions.
Definition
CO2-fixation reactions
Term
What is the primary reaction by which CO2 is fixed in most plants? What enzyme is the reaction catalyzed by?
Hint: Ribulose
Definition
Bisphosphate + CO2 --> 2 3-phosphohulcerate
Enzyme: ribulose bisphosphate carbozylase (or RuBP carboxylase)
Term
RuBP carboxylase often accounts for more than ___% of the soluble protein in leaves.
Definition
50
Term
Photosynthesis consists not only of this series of physical and chemical reactions occurring in the chloroplasts, but also of the ______________ between the leaf and its atmospheric environment.
Definition
exchange of matter
Term
Describe how CO2 reaches the site of carboxylation after it diffuses from the atmosphere to the leaf surface.
Definition
It goes through the stomata, across the intercellular air space, into the cell, across the cytoplasm, and into the chloroplast, where it finally reaches the site of carboxylation.
Term
____________ is a spontaneous process leading to the net movement of a substance from a region of higher concentration to an adjacent region where that substance has a lower concentration.
Definition
diffusion
Term
In the case of CO2, its concentration in the atmosphere is usually much ___________ than its concentration at the site of carboxylation in the chloroplast.
Definition
higher
Term
A key issue for plants living in dry environments is _________________.
Definition
photosynthetic water-use efficiency
Term
What is the equation for photosynthetic water-use efficiency? What are the units?
Definition
amount of CO2 fixed by photosynthesis/amount of H2O lost by transpiration

unit: mol
Term
In most plants, the primary fixation product of photosynthesis is the three-carbon compound
_______________ (C3).

However, in some plants, the primary fixation product of photosynthesis is the four-carbon compound _____________ (C4).
Definition
3-phosphoglycerate; oxaloacetate
Term
In C4 plants, what is the primary reaction by which CO2 is fixed? What enzyme catalyzes the reaction?
Definition
Reaction: Phosphoenol pyruvate + CO2 -> oxaloacetate
Enzyme: phosphoenol pyruvate carboxylase (or PEP carboxylase)
Term
The primary reaction of C4 plants occurs in specialized __________ cells in the leaves.
Definition
mesophyll
Term
Oxaloacetate is converted into malate, which is transported into specialized ___________ cells.
Definition
bundle sheath
Term
What are the two special properties of PEP carboxylase relative to RuBP carboxylase?
Definition
1. PEP carboxylase has a much higher affinity for CO2 than does RuBP carboxylase.

2. PEP carboxylase is not competitively inhibited by O2, whereas RuBP carboxylase is.
Term
C4 plants can maintain _________ rates of photosynthesis with lower degrees of stomatal opening than C3 plants.
Definition
higher
Term
C4 plants have intrinsically _________ photosynthetic water-use efficiencies than C3 plants.
Definition
higher
Term
C4 plants are most abundant in ______ environments where photosynthetic water-use efficiency is at a premium.
Definition
dry
Term
A third major biochemical pathway of photosynthesis, known as ___________________, is found in other plants.
Definition
CAM photosynthesis
Term
CAM stands for __________________, so named because it was originally found in the plant family Crassulaceae.
Definition
Crassulacean Acid Metabolism
Term
CAM photosynthesis is now known to occur in about 20 plant families, including those with species such as ______ cacti, _________ cacti, _______ cacti, and ______ from the desert surrounding Tucson.
Definition
saguaro; prickly-pear; barrel; agaves
Term
CAM plants are ___________ (or fleshy), with the leaf or stem cells containing large __________.
Definition
succulent; vacuoles
Term
Meiosis always gives rise to _________ products.
Definition
haploid
Term
CAM plants exhibit a _______________, in which the stomata open during the night and close during the day.
Definition
reversed stomatal cycle
Term
When the stomata are open at __________, CO2 is fixed by PEP carboxylase into oxaloacetate, via the same reaction as in C4 plants.

_____________ is then converted into malate, which is stored in the large vacuoles.
Definition
night; oxaloacetate
Term
During the ______, when the stomata are _______, the malate is removed from the large vacuoles and decarboxylated, with the released CO2 then being fixed by RuBP carboxylase via a pathway similar to that in C3 plants.
Definition
day; closed
Term
In CAM plants, the reactions catalyzed by PEP carboxylase and RuBP carboxylase occur within the ________ cells.

This contrasts with the situation in C4 plants, where the reactions catalyzed by PEP carboxylase and RuBP carboxylase occur in _________ cells.
Definition
same; different
Term
The photosynthetic reactions involving PEP carboxylase and RuBP carboxylase are separated in ________ in CAM plants, whereas they are separated in _______ in C4 plants
Definition
time; space
Term
By opening their stomata at night and closing them during the day, CAM plants are able to achieve very ________ photosynthetic water-use efficiencies, typically much _________ than those of C4 plants.
Definition
high; higher
Term
CAM plants usually exhibit ______ photosynthetic rates, and thus typically have ______ growth rates.
Definition
low; low
Term
_____________ is a perennial shrub that remains physiologically active throughout the year, even though air temperatures vary greatly from winter to summer.
Definition
Creosote bush
Term
Plants growing under cool winter conditions exhibited ________ photosynthetic rates at ______ leaf temperatures, whereas plants growing under hot summer conditions exhibited _________ photosynthetic rates at ________ leaf temperatures.
Definition
high; low; high; high
Term
What did Stanford biologists find when they analyzed the biochemical properties properties of leaves of creosote bush and other species?
Definition
1. leaves of plants growing under cool winter conditions had higher levels of the enzymes fructose-phosphate phosphatase and RuBP carboxylase than leaves of plants growing under hot summer conditions, and

2. leaves of plants growing under hot summer conditions exhibited an increase in the thermal stability of the reactions associated with the thylakoid membranes in the chloroplasts relative to leaves of plants growing under cool winter conditions.
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