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Evolution Exam 2
Exam 2 Evolution
98
Biology
Undergraduate 4
11/06/2012

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Term
What helped lead Darwin to his theory of why species changed?
Definition
Malthus’ essay on limits to population growth and struggle
Term
Because of the struggle for existence, forms that are better adapted to survive will leave more....
Definition
offspring and automatically increase in frequency from one generation to the next
Term
What is natural selection the logical result of?
Definition
Superfecundity
Variation
Heritability
Term
Why do organisms compete for survival?
Definition
Organisms produce more offspring than - given the limited amounts of resources - can ever survive
Term
What does the struggle for existence take place within?
Definition
a web of ecological relations
Term
Why does an organism compete most closely with other members of its own species?
Definition
because they have the most similar ecological needs to its own.
Term
In every species, more eggs are produced than can survive to the adult stage
Why does excess fecundity exist?
Definition
because there are inadequate resources in nature to support all the eggs that are laid and all the young that are born
Term
What provides the preconditions for natural selection?
Definition
The excess fecundity, and consequent competition to survive in every species
Term
Review Slide 6 in Presentation 7
Definition
Term
What are the 4 conditions for natural selection to occur?
Definition
Reproduction

Variation in individual characters among the members of the population.

Heritability

Variation in the fitness of organisms according to the state they have for a heritable character.
Term
What is heritability (broad definition)?
Definition
the proportion of variation (or variance) in a phenotypic character in a population that is due to individual differences in genotypes.
Term
What is heritability (narrow definition)?
Definition
the proportion of variation (or variance) in a phenotypic character in a population that is due to individual genetic differences that will be inherited in the offspring
Term
When does natural selection occur?
Definition
When we have differential survival and reproduction.
Term
Describe what it means to have higher fitness? Describe the process.
Definition
- individuals in the population with some characters must be more likely to survive and reproduce

- The average number of offspring left by an individual with a certain genotype relative to the number of offspring left by an average member of the population or relative to the number produced by individuals of other genotypes.
Term
In natural selection, why does incidental selection of other features occur?
Definition
Because they are correlated with the fatures that are being selected.
Term
Review Slide 12, Class 7. IT'S INTERESTING!
Definition
Term
What is the definition of adaptation?
Definition
a phenotypic variant that results in the highest fitness among a specified set of variants in a given environment
Term
For a character to be regarded as an adaptation, it must be a...
Definition
derived character that evolved in response to a specific selective agent
Term
What is Exaptation?
Definition
the evolution of a function of a character other than the function the character originally adapted to fulfill.
Term
What is an adaptation?
Definition
a
phenotypic variant that results in the highest fitness among
a specified set of variants in a given
environment
Term
What does directional selection cause?
Definition
causes a consistent directional change in the form of the population through time.
Term
What does Stabilizing selection tend to do?
Definition
keep the form of the population constant.
Term
What does disruptive selection do?
Definition
favors forms that deviate in either direction from the population average.
Term
What are the four types of selection?
Definition
Disruptive, directional, stabilizing, no selection
Term
What are the two different types of variation in biology?
Definition
-characters of an organism which can vary among individuals at both the phenotypic and the genetic levels.

-variation in the fitness of individuals.
Term
What are the 5 levels of variation in nature?
Definition
The morphologic level
The physiological level
The cellular level
The biochemical level
The DNA level
Term
What are some sources of genetic variation in a population?
Definition
mutation, recombination, migration
Term
What is the "original" source of genetic variation?
Definition
mutation
Term
? between chromosomes produces new chromosomes.
Definition
Recombination
Term
if there is no ?, recombination and migration would not generate new genetic variants.
Definition
pre-existing allelic variation
Term
What are the 4 types of genetic mutations?
Definition
synonymous mutation
amino acid changing mutation
frame-shift mutation
stop mutation
Term
Mutations do not arise in the direction of ?
Definition
improved adaptation
Term
Filter variation by natural selection to produce adaptations: random or not random?
Definition
not random
Term
Natural selection is the ? survival of ? variants.
Definition
non-random; random
Term
5 Factors affecting the distribution of variation in natural populations
Definition
Natural Selection
Mutation
Genetic Drift
Inbreeding
Migration
Term
When does random sampling occur?
Definition
whenever a smaller number of successful individuals (or gametes) are sampled from a larger pool of potential survivors and the fitnesses of the genotypes are the same.
Term
What kind of important evolutionary effects can random sampling have?
Definition
genetic drift and the founder effect
Term
Genes that form a new generation are a ? from the parental generation
Definition
random sample
Term
The frequency of alleles with the same fitness will ? through time
Definition
change at random
Term
If there are two alleles at a locus, and they have the same fitness, random sampling can cause their...
Definition
relative frequencies in a population to change.
Term
What is genetic drift?
Definition
Random changes for a gene in allele frequencies between generations
Term
What does the rate of change of gene frequency by random drift depend on?
Definition
the size of the population
Term
Are random sampling effects more important in a bigger or smaller population?
Definition
smaller
Term
What is a locus?
Definition
the specific location of a gene or DNA sequence on a chromosome
Term
The Hardy-Weinberg ratios are for neutral alleles at a locus and the Hardy-Weinberg result suggested that...
Definition
the genotype (and gene) ratios are stable over time.
Term
In a small population, gene frequencies drift about; one of the genes will eventually be fixed: and only then will...
Definition
the system be stable.
Term
In a small population, gene frequencies drift about; one of the genes will eventually be fixed: and only then will...
Definition
the system be stable.
Term
What is the true equilibrium of the Hardy-Weinberg system in a small population?
Definition
homozygosity
Term
Over the long term, in Hardy Weinberg Ratios, what causes the population to "march" towards homozygosity (decline in heterozygosity) at a locus?
Definition
pure random drift
Term
The rate of decline in heterozygosity is often used as a measure of what?
Definition
the rate of genetic drift within a population.
Term
The rate of decline in heterozygosity is often used as a measure of the rate of genetic drift within a population.
If this process occurs in a large number of independent, non-interbreeding populations, the genetic composition of the populations would...?
Definition
diverge by chance.
Term
At any time, what does an allele’s probability of fixation equal? and what is it not affected by?
Definition
its frequency at that time;its previous history of change in frequency.
Term
The probability is finite, however, and if enough neutral alleles, at enough loci, and over enough generations, are randomly drifting in frequency, one of them will eventually..
Definition
be fixed
Term
For any allele, is fixation by random drift very probable or improbable?
Definition
improbable
Term
What is the founder effect? Who is credited with describing the founder effect?
Definition
“The establishment of a new population by a few original founders (in an extreme case, by a single fertilized female) which carry only a small fraction of the total genetic variation of the parental population.”; Ernst Meyer
Term
What is the he founding effect quite ineffective at?
Definition
reducing genetic variation.
Term
What is the most likely explantation for isolated populations often having exceptionally high frequencies of otherwise rare alleles?
Definition
the founding population had a disproportionate number of those rare alleles.
Term
Give an example of the founder effect and explain why it is an example of that?
Definition
European Bison- last one killed in the wild in 1927, but 50 still survived in zoos. Today's population of approximately 3,000 has descended from about 12 individuals
Term
Describe the founder effect being evident in amish communities
Definition
Ellis-van Creveld syndrome, involves not only short stature but polydactyly (extra fingers or toes), abnormalities of the nails and teeth, and, in about half of individuals, a hole between the two upper chambers of the heart.

It has been traced back to one couple, Samuel King and his wife, who came to the area in 1744.
Term
Natural selection will act to eliminate any allele that decreases the ...?
Definition
fitness of its bearers
Term
What is recurrent disadvantageous mutation?
Definition
Selection can never finally eliminate the gene, because it will keep on reappearing by mutation.
Term
In recurrent disadvantageous mutation, what is the equilibrium between?
Definition
the mutant gene's creation, by recurrent mutation, and its elimination by natural selection.
Term
What does the theory of neutral drift predict?
Definition
populations should be completely homozygous
Term
Give two examples of reduced polymorphism because of genetic drift.
Definition
By 1900 hunting of the northern elephant seal off the Pacific coast had reduced its population to only 20 survivors. The population today exceeds 100,000. However, these animals are homozygous at every one of genes that has been examined.

The lack of genetic variability in cheetahs is so profound that cheetahs will accept skin grafts from each other. 52/52 genes tested showed no polymorphisms.
Term
What is inbreeding?
Definition
a form of nonrandom mating, which occurs when individuals are more likely to mate with relatives than nonrelatives.
Term
With inbreeding in a finite population, ? is increased and ? is decreased over the Hardy-Weinberg level generated by random mating even though the allelic frequencies do not change each generation.
Definition
homozygosity; heterozygosity
Term
The important point is that inbreeding can happen in any breeding system with a small population, and becomes more likely the smaller or larger? the population
Definition
smaller
Term
Example of inbreeding?
Definition
The great tit- the less eggs hatch the more likely it is to inbreed because of less members in a population
Term
What is migration?
Definition
Gene flow between populations is due to movement of individuals from one population to another.
Term
If unopposed by other factors, what does gene flow do?
Definition
homogenizes the populations of a species or brings all populations to the same allele frequencies.
Term
What happens in the absence of gene flow or low gene flow between populations?
Definition
allele frequencies will differ among populations.
Term
Genetic difference among populations typically correlates with...?
Definition
geographic distance
Term
What is effective population size?
Definition
the number of individuals in a population who contribute offspring to the next generation.
Term
Factors affecting the distribution of variation in natural populations?
Definition
Natural Selection
Mutation
Genetic Drift
Inbreeding
Migration
Term
Any factor which increases the chance that copies of the same gene will combine in the same individual in the production of the next generation will lower what?
Definition
the effective population size relative to the total number of adults
Term
What are the factors affecting the distribution of variation in natural populations?
Definition
Natural Selection
Mutation
Genetic Drift
Inbreeding
Migration
Term
In response to selection on phenotype, genotype frequency...
Definition
increases or decreases
Term
Selection only works directly on ? not ?
Definition
phenotype; genotype
Term
Selection only works directly on phenotype not genotype, which reduces or increases genotype frequency only as far as...
Definition
how close the one to one match is between genotype and phenotype
Term
The response to selection or evolution for polygenic traits (e.g., beak thickness) depends on the
Definition
heritability of the trait
Term
high narrow-sense heritability then have a large amount of
Definition
additive genetic variability
Term
What does directional selection reduce additive genetic variability with ?
Definition
time
Term
When ? are fitter than the ?, therefore, natural selection will maintain a polymorphism.
Definition
heterozygotes; homozygotes
Term
What is polymorphism?
Definition
having multiple phenotypes within a population
Term
What is one of several possible explanations for the existence of genetic variability in natural populations?
Definition
Heterozygous advantage
Term
What is an example of heterozygous advantage?
Definition
sickle cell anemia in areas of malaria in Africa.
Term
When does frequency-dependent selection occur?
Definition
when the fitness of a genotype depends on its frequency. It is possible for the fitness of a genotype to increase (positively frequency-dependent) or decrease (negatively frequency-dependent) as the genotype frequency in the population increases.
Term
What is a Mendelian Population?
Definition
a group of interbreeding, sexually reproducing individuals with a common set of genes: the gene pool.
Term
The genetic composition of a population can be described in terms of what?
Definition
frequencies, or relative abundances, in which alternative alleles are found.
Term
Genotypes differ in ? from one population to another.
Definition
frequency
Term
What is population genetics concerned with?
Definition
gene frequencies and genotype frequencies, and with the processes influencing these frequencies.
Term
What is the most important, most fundamental body of theory in evolutionary biology?
Definition
Population genetics
Term
The coherence of an evolutionary hypothesis will usually remain in doubt until...?
Definition
the hypothesis is expressed in the form of a population genetic model.
Term
What does allele frequency measure?
Definition
the frequency in the population of a particular allele relative to other alleles at one gene locus.
Term
Why is H-W an equilibrium?
Definition
because the new frequency of A or p’ is the same as the old frequency of A or p
Term
What are the assumptions for the Hardy-Weinberg equilibrium?
Definition
Large population and random mating
No mutation, migration, and natural selection
Term
What are the predictions for the Hardy-Weinberg equilibrium?
Definition
Allelic frequencies remain constant from generation to generation
2. Genotypic frequencies stabilize (will not change) after one generation in the proportions
HWE = AA: p2 Aa: 2pq aa:q2
Term
Three sources of interest in the Hardy-Weinberg ratio?
Definition
Historical and Conceptual interest
Deviations direct future research
Theoretical interest
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