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the study of the distribution and abundance of organisms and the factors and interactions that determine distribution and abundance |
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Historia Animalium: patterns of the natural world explained by natural causes |
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Providential ecology: balance of nature |
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father of demography: study of population growth |
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how populations grow over time |
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how predators regulate prey population: population regulation |
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used exponential growth to explain revolutions |
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Quetelet 1835, Verhulst 1838 |
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proposed and studied the idea of logistic population growth |
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logistics curve: population regulation |
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Introduced a rule regarding the relationship between population size and the death rate: as density increases mortality increases. |
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Edward Forbes 1887 HC Cowles 1899 |
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community regulation and succession, mostly looked at patterns |
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systems analysis mathematical model of the spread of infectious disease (malaria) |
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AG Tansley 1904 FE Clements 1905 Charles Elton 1927 |
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some of the founders of modern ecology (experimental): why pattern occured |
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helped make ecology accepted by the public- wrote Silent Spring, a work exposing the negative effects of pesticide use |
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non-living chemical and physical factors |
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Does ecology typically operate at the high of low scales of biological organization? |
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a single individual or a single species |
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individuals of the same species living in the same geographical area |
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2 or more populations living in the same geographical area |
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comprising of the community and its physical environment |
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lithosphere (rock), hydrosphere (water) and atmosphere (gas) |
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p-values: measure the strength of conclusions being drawn Null hypothesis: assume there is no association between variables Significance testing: if P is less than 0.05, then the results are statistically significant |
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If a species is in a certain area... |
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1. it evolved there 2. it evolved elsewhere and the dispersed to that area |
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If a species is not found in a certain area... |
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1. it evolved elsewhere and never dispersed there 2. it was once present but no longer is there |
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Based on the taxonomic similarities of the organisms living there (6 major regions) Boundaries are set where species composition changes dramatically over short distances |
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the short term state of atmospheric conditions at a particular place and time |
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the long term average atmospheric conditions found over time (temperature, wind velocity, precipitation) Climate varies because of differences in the amount of solar energy (drives global atmospheric and oceanic circulation) Large scale distribution patterns can be predicted from climate conditions |
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rotation of Earth on its axis 24 hour cycle |
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Seasonal Temperature Cycle |
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annual cycle- fixed axial tilt at 23.5 degrees variation in sunlight intensity and day length give rise to seasonal variation in temperature in north and south hemispheres |
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Why are the tropics hotter than temperate regions? (3) |
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1. solar radiation strikes at almost right angles near the equator but at acute angles near the poles 2. same amount of radiant heat is concentrated over a smaller area at the tropics 3. at poles solar radiation passes through a deep layer of atmosphere (reflection, absorption, scattering) |
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Global Air Circulation Pattern (4) |
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1. Wet hot air: radiant heat above equator warms the air (low pressure) at equator- moist, hot air rises to cooler altitudes and rains (tropical rainforests) 2. Hot dry air: cool air descends at latitudes of about 30 degrees north and 30 degrees south and heats (deserts) 3. Wet cool air: air picks up moisture and rises at 60 degrees north and south and rains (temperature rainforests) 4. dry cool air: drier air descents in polar regions (cold dry polar deserts) |
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What determines the distribution of biomes? |
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What are terrestrial biomes based on? |
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The structure of their dominant vegetation. In each terrestrial biome vegetation has similar traits, but may not be closely related- looks similar and functions in a similar way but species could be totally different |
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What are aquatic biomes based on? |
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Their physical and chemical differences |
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how to analyze a distribution on a smaller scale- see what factors are limiting the distribution. move organism and see if it can survive and reproduce in a new environment- must go through at least one generation |
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If a transplant is successful... |
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potential range of a species is larger than its actual range. Lacks means of transport (dispersal) Can move but chose not to (habitat selection) |
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If a transplant is unsuccessful... |
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potential range of a species is the same as its actual range. limits imposed by other species (predators, competitors, parasites, disease) positive effects of interdependent species or some other physical/chemical barrier (too hot, too cold, etc.) |
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Factors Limiting Distribution: Dispersal |
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the movement of organisms from one geographical location to another. most organisms are mobile geographical barriers are common in nature. barrier to dispersal- breached by human introduction. have to control for the experiemental factors |
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Factors Limiting Distribution: Habitat Selection |
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organisms can move but choose to not live in certain habitats. distribution may be limited by the behavior of individuals in selecting their habitat behaviorally constrained distribution |
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Factors Limiting Distribution: Competition |
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-,- interaction: both species in the interaction have negative consequences. can occur between 2 or more organisms or species that exploit the same types of limited resources and live in the same geographical area Interference competition: direct physical interaction over resources- large consequences Exploitative competition: indirect interaction over resources |
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Natural patterns indicating competition |
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1. When species B is absent, species A lives in a wider range of habitats- Competitive release 2. if competition is strong, the geographic ranges of the two competitors may not overlap but have sharp boundaries |
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Factors Limiting Distribution: Predation |
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+,- interaction Predation can limit the distribution of organisms via: 1. direct consumption of the prey 2. behavioral modifications of prey in the presence of predators |
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Factors Limiting Distribution: Physical and Chemical |
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temperature and availability of water are the 2 major limiting factors to the distribution of life on earth. organisms have upper and lower lethal limits to physical and chemical factors. Solutions are acclimation (dealing with it short term) or adaptation (evolutionary changes). Also light and soil structure limits distributions |
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light is important to the development and behavior of many plants and animals that are sensitive to this period- drives the distribution of plants and thus animals |
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Soil structure as a limiting factor |
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the composition of substrate affects water chemistry of streams and rivers which influences the resident plants and animals
Ex: serpentine soil is high in nickel and magnesium- only Goldfield flowers can prosper because they use magnesium instead or calcium. Goldfield flowers can survive in other soils but they are poor competitors |
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Our ecosystem in SB, fire may be unpredictable in the short term but predictable in the long term so many plant communities have become adapted to fire and are dependent on fire to maintain them- ash is a major fertilizer, fire opens up space and light |
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Factors Limiting Distribution: Disturbance |
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fires, hurricanes, tornadoes, and volcanoes Frequent disturbance: may be predictable and organisms may be evolutionarily adapted to the disturbance Infrequent disturbance: may be unpredictable and organisms will not have acquired evolutionary adaptations to them- most damaging |
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Factors Limiting Distribution: pH and Salinity |
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affects the distribution of marine and freshwater organisms |
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Factors Limiting Distribution: Currents, Oxygen |
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many organisms dependent on wind and water currents to disperse their propagules or young oxygen can limit the distribution or organisms in lakes important in aquatic environments |
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Characteristics of Populations |
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Natality, Mortality, Immigration, Emigration |
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the number of individuals per unit area (terrestrial) or volume (aquatic) |
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How do you measure population density? (3) |
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1. Total counts (almost impossible) 2. Sub-sampling methods: sub sample population to estimate densities and total population size (big assumption) 3. Indirect Indicators: number of nests, fecal droppings, tracks, etc. as estimates of density |
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Patterns of Dispersion (3) |
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1. Clumped: most common, due to unequal distribution of resources and/or social behavior 2. Uniform: distributed based on minimum distance between indiv., due to interactions between individuals 3. Random: (least common) distributed without regard to position of other individuals |
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Are emigration and immigration more important to small or large spatial/temporal scales? |
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The smaller the spatial/temporal scale, the greater the importance of movement to population dynamics |
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Methods used to identify how population size changes (3) |
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1. Direct observations: not practical due to long lived organisms 2. View age structures: - Type 1: humans, live for a long time and then die -Type 2: birds, die at same percent per unit time -Type 3: plants and fish, when you are born you have a high chance of death but then live for a very long time 3. Mathematical models |
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maximum population size that an environment can support population growth slows as its density approaches |
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Why does the logistic curve not fit well for most natural population growth patterns? (name assumptions, 3) |
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1. Each individual added to the population has the same negative effect on population growth- less resources (exceptions: isolated plants and flamingos) 2. Population approaches K smoothly: lag time causes the population to overshoot and undershoot, oscillate about K 3. populations are large and density is important in regulation |
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Density Dependent Control (limiting factor) |
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factors that alter per capita birth or death rates in a population are dependent on population density (can stabilize population density) -as density goes up, higher death rate -bubonic plague and fleas: spread disease because they kept biting -resources -competition predation |
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Density Independent Control (limiting factor) |
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factors that alter per capita birth or death rates in a population are independent of population density (do not stabilize population density) -percent death does not change no matter what the population density is -monarch butterfly- freeze wiped them out, didn't matter how many there were -usually weather and disturbances |
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Are short term fluctuations in populations due to density dependent or density independent factors? |
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density- independent factors |
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if perturbed population will return to initial density stabilizing forces dampen population fluctuation density dependent controls birth= death |
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if perturbed, population may not return to initial density destabilizing forces enhance population fluctuations inverse density dependence- density dependence with long time lags |
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populations is divided into discrete sub-populations connected by immigration and emigration fast increase in migration: local population fluctuations are damaged migration decrease: local population fluctuations are enhanced but there is an increased probability of extinction intermediate migration: how most of the world works: shifting mosaic or occupied and unoccupied patches |
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+,+ both species benefit from the association ex: large African mammals and tick birds |
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+,0 one species benefits from the association, the other is unaffected (it doesn't affect the population) ex: remora and shark |
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-,0 one is unaffected and one has a negative affect ex: walnut tree shading smaller trees |
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What are the possible outcomes of competition? (3) |
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1. both species coexist 2. species 1 becomes extinct 3. species 2 becomes extinct |
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Competitive Exclusion Principle (Gause Hypothesis) |
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No two species can occupy the same ecological niche simultaneously (complete competitors cannot exist) |
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the biological role of an organism within the environment, where the organism fits in to |
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Fundamental Niche (n dimensional hypervolume) |
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the multitude of conditions in which an organism can survive and multiply- the space you can potentially occupy |
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where an organism actually exists due to ecological constraints |
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Where does competitive exclusion not occur? (3) |
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1. Unstable environments: never reach equilibrium, low density of organisms- population sizes aren't big enough to dominate resources 2. Environments in which species do not compete (unlimited resources or partitioning- giraffes eat higher up, zebras in the middle) 3. Fluctuating environments that reverse the direction of competition before extinction occurs- system changes so competitors never become too successful |
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Carnivory
Parasitism
parasitoids
cannibalism
herbivory |
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Lotka-Volterra Predator Prey Model Assumptions (2) |
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1. In the absence of predators, prey populations grow exponentially- assume unlimited resources
2. In the absence of food, predator populations decline exponentially
These predator prey populations oscillate, but few good examples in natural populations |
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1. prey have a spatial (hiding) or temporal (eating at different times) refuge
2. optimal foraging: predators switch to other prey as original prey species fall to low abundance
3. antipredator strategies:
-chemical defense: aposematic coloration (to warn) and poison
-camouflage: cryptic coloration and mimicry
-displays of intimidation and fighting
-agility
-armor
-altered reproductive patterns: masting, so many offspring you overwhelm predator |
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Individualistic Hypothesis |
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Gleason
communities are chance assemblages of species with similar abiolic requirements
a more general way to say nonequilibrium view |
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Clements
communities are assemblages of dependent closely linked species
a general way to say equilibrium view |
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Features of ecosystems (3) |
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1. open biological systems
2. one way flow of energy
3. cycling of nutrients: energy imput from the sun is made into nutrients |
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group species by functional similarity
defined by the number of steps through which energy passes to reach the organisms in it
1. primary producers
2. primary consumers: herbivores, decomposers, detritivores (eat primary producers)
3.secondary consumers (eat primary consumers)
4. tertiary consumers (eat secondary consumers)
5. quarternary consumers (eat tertiary consumers)
forms a hierarchy of feeding relationships- shows flow of energy through ecosystem
most communities only have 3-5 trophic levels |
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measure of weight of dry tissue (carbon) an organism contains |
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An average of ___ % of energy is transferred from one level to the next |
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the abundance of lower trophic levels depends on the effects of consumers from higher trophic levels |
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the abundance of higher trophic levels depends on factors such as nutrients and prey availability from lower trophic levels |
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Odd number of trophic levels means...
Even number of trophic levels means... |
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odd- resources limitation
even- consumer limitation
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species richness + species evenness |
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the number of species in a community |
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the relative abundance of a species in a community |
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Does more diversity mean more or less stability? |
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an event that removes organisms and alters resource availability
communities with very high or low disturbance have lower species diversity than communities with moderate disturbance |
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Intermediate disturbance hypothesis |
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1. when disturbance is severe and frequent community is composed of good colonizers (r-selected) with high reproductive rates
2. when disturbance is mild and rare community is composed of good competitors (k-selectors)
Species diversity is highest where disturbance is moderate (intermediate) in severity and frequency- allows both colonizers (r) and competitors to coexist (k) |
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The Diversity-Stability Hypothesis |
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1. disturbance in diverse communities is dampened by large numbers of interacting species
2. the effect of disturbance is less than it would be in a species poor communities
Higher diversity increases community stability |
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Transition in species composition over time
Primary succession: bare soil, no organisms
Secondary succession: soil, some organisms |
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1. Early successional communities (r-selection): good dispersers (colonizers) and high reproductive rates
2. Late successional communities (k-selection): good competitors, exist near carrying capacity |
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Successional patterns (2) |
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1. Directional succession: progressive change in species composition
2. cyclical succession: change in species composition in which the original community is ultimately restored |
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Biological invasion and impacts |
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Invasive species: species that disrupt communities by a dominant colonization of a particular habitat or from loss of natural controls (predators or herbivors)
Impacts: changes in ecosystem structure and function
Environmental disturbance promotes biological invasion which can alter native succession and promote more invasion |
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the wider the average niche breadth the fewer the number of species in the community (extinction via competitive exclusion)
the narrower the average niche breadth the more species in the community (coexistence via resource partioning) |
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How can predation increase richness? |
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Predation may increase richness by allowing compeitively inferior species to coexist with superior competitors
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a species influence on the community is greater than would be expected on the basis of abundance
moderate levels of predation increase species diversity |
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Equilibrium model of island biogeography |
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species richness is positively correlated with island size and negatively correlated with distance from the mainland
if extinction is greater than immigration then species diversity decrease
if extinction is less than immigration then species diversity increases |
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1. communities are structured by biotic interactions (competition, predation, herbivory)
2. communities display global stability
3. niche diversification determines species diversity
interactive hypothesis |
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1. communities composition is always changing and never in balance
2. no global stability
3. communities are patchworks of disturbance. Colonization and dispersal determine species diversity
individual hypothesis |
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1. genetic diversity (amount of genetic variation among species)
2. species diversity (number and relative abundance of species)
3. ecosystem diversity (diversity of structure and function in an ecosystem)
Conserve biodiversity because of economic value, ecosystem services, and environmental ethics |
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Inverse Density Dependence |
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each individual added actually increases the population
trees: more of them, work together to bring up the water
flamingos |
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