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Bryophytes vs. Vascular Plants |
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-Bryophytes: dominant gametophyte (n), lg.er than sporophyte, liverwort -Vascular Plants: xylem & phloem, lignin, branched & independent sporophyte, dominant sporophyte (2n), multiple sporangia produced on one plant, free-living sporophyte |
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Pivotal Steps in the Evolution of Plants |
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-vascular tissue: plants can grow tall, & able to support very negative turgor pressure -lack of reliance on water for sexual reproduction -evolution of leaves: no true leaves in mosses |
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-must have vascular tissue to be considered a leaf -2 types: Microphylls & Megaphylls |
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-sm.er than megaphylls -have just 1 strand of vascular tissue -1 vein -vascularization of enations |
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-lg.er than microphylls -have multiple vascular bundles -more than 1 vein -dichotomous branching -unequal branching -planation of branching -webbing |
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-lycophytes -herbaceous -have microphylls -3 families: Lycopodiaceae, Selaginellaceae, & Isoetaceae |
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-family in Lycopodiophyta -branched sporophyte |
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-cones -areas where sporophylls packed together |
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-family in Lycopodiophyta -heterosporous: 2 types of spores -2 types: microspores & megaspores |
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-Microsporangium -Microsporophyll -Microgametophyte: produces sperm |
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-Megasporangium -Megasporophyll -Megagametophyte: produces eggs |
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-family in Lycopodiophyta -quillworts -heterosporous -aquatic -secondary growth |
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-ferns: Eusporangiate & Leptosporangiate -4 Orders: Order Filicales, Order Salviniales, Order Psilotales, & Order Equisetales |
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-homosporous -Leptosporangiate |
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-cluster of sporangia on abaxial surface of leaves |
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-protective cap covering sorus |
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-heterosporous -Leptosporangiate |
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-genus in Order Psilotales -leafless, but has enations -dichotomous branching |
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-genus in Order Equisetales -horsetails -homosporous -produce sm. strobili -microphylls fused into sheaths -silica in cells |
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-broad leaves & spores -sm. & non-woody |
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-diploid, haploid, & triploid -embryo |
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-produced on surface of leaves in sporangia -disperse via wind -grows into gametophyte (n) -heterosporous: microspores & megaspores |
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In Seed Plants (Spermatophytes) |
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Definition
-megaspores retained in megasporangium (not released) -reduction in megaspore mother cell = cells that produce spores = sm.er # of spores produced -only 1 megaspore produced per megasporangium -megagametophyte grows inside spore wall -formation of integument/strong tissue, surrounding megasporangium -embryo develops inside the megaspore -micropyle & ovule |
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-one opening in integument for sperm |
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-integuments -megasporangium -megaspore -develops into seeds & disperses |
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-paraphyletic group -"naked" seed b/c seeds produced on surface of sporophylls -contains cycads, ginkgos, conifers, & gnetales |
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-conifers & "softwoods" -most widespread & economically important phylum of gymnosperms -all tracheids in vascular tissue, no vessel elements -genus Pinus -Pollen Cone/Microsporangiate strobilus -Ovulate Cone |
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-pines (produce seeds) -in family Pinaceae & Phylum Coniferophyta -leaves "needle-like" -leaves usually grouped in clusters -fascicles |
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Pollen Cone/Microsporangiate strobilus |
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-microsporophylls -1-2 cm long -on lower branches of trees -each microsporangium contains many microspore mother cells -undergo meiosis to produce microspores -develops air bladders & 4 internal cells: pollen |
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-made up of seed-scale complexes |
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-most look like palm trees -Dioecious -250 million yrs ago in fossil record -abundant food source for dinosaurs -in roots: house cyanobacteria & produce toxic compounds |
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-2 houses -"male" and "female" individuals |
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-1 family: Ginkgoaceae -1 genus & 1 species: Ginkgo biloba -fan-shaped leaves & veins dichotomously branch -Dioecious -maintained throughout history as horticultural specimen in Chinese & Japanese temples |
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-3 families: Gnetaceae, Ephedraceae, & Welswitschiaceae |
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-in Phylum Gnetophyta -Genus Gnetum -produces broad leaves that resemble leaves of flowering plants -thought to be closely related to flowering plants -produces vessel elements -sporophylls may resemble those in flowering plants |
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-in Phylum Gnetophyta -Genus Ephedra -grows in SW U.S.A. -produces ephedrine (chemical) |
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-in Phylum Gnetophyta -Genus Welswitschia -grows in sand dunes in Namibia |
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-lg.est phylum of photosynthetic organisms -angiosperms: flowering plants, "closed seed" -woody or herbaceous -fruits produced -seeds |
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-determinate shoot -growth for limited duration -appendages -4 floral whorls: sepals, petals, stamens, & carpels -perianth -tepals |
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-sporophylls (sporangium-bearing leaves) -sterile appendages (leaf-like, but not sporangia) |
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-outermost whorl -usually green/leaf-like -calyx: all sepals together |
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-usually brightly colored & lg.er than sepals -corolla: collectively |
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-variety of sizes/shapes -anther: microspangia & produce pollen -adroecium: male house & stamens all together |
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-in center of flower -megasporophylls: produce megasporangia -usually fused together into one or more pistil(s) -ovary contains ovules: produce eggs -gynoecium: women house & all carpels together |
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-sepals & petals together -not usually containing sporangia |
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-when sepals & petals indistinguishable |
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-perfect flower -stamens & carpels |
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-either only carpels or only stamens, or if one is nonfunctional -imperfect flower |
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-one or more whorls missing |
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-fusion among members of the same whorl -ex: all petals fused |
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-fusion among members of different whorls -ex: stamens fused to petals |
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-superior ovary: sepals, petals, & stamens attached below ovary -inferior ovary: sepals, petals, & stamens attached above ovary |
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-one single flower on stem |
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-group of flowers together -head, spike, & catkin |
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-flowers packed together in flat disk |
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-1 shoot w/flowers grouped close together |
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-chain of flowers that hang -slender chain of sm flowers that's produced in some wind-pollinated trees -sepals and petals sm or absent -no nectar |
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-contains microsporangia -microsporophytes -spores develop into pollen (immature male gametophyte) -sporopollenin & pollen |
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-2n cells that undergo meiosis to produce haploid microspores |
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-strong polymer -outer layer of pollen |
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-inside: 2 cells (tube cell & generative cell) -transported through some vector through space -to stigmatic surface |
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-may be able to sense if pollen is from same species -pollen produces pollen tube that grows down the style -1 sperm fertilizes egg in ovule & other sperm combines w/polar nuclei to form 3n structure |
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Integuments to Egg to Polar Nuclei to Ovule to Ovary |
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-seed coat -embryo -endosperm -seed -fruit w/seeds |
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-only in Angiosperms -following reproduction, ovaries turn into fruit & ovules turn into seeds -primarily used for seed dispersal -fleshy: may attract animals -winged fruit: facilitates wind dispersal of seeds |
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-known as Pericarp -endocarp, mesocarp, & exocarp make up the pericarp -endocarp is inside near the seeds, & exocarp is towards the outside |
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-produced from 1 ovary in 1 flower -5 types: berry, hesperidum, drupe, coir, & pepo |
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-simple fruit w/fleshy mesocarp -ex: tomato, banana, & bell pepper |
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-simple fruit w/leathery exocarp -ex: citrus fruits |
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-simple fruit, fleshy, endocarp hard & fused to seed -ex: olive, avocado, & coconut |
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-simple fruit w/fibrous material that makes up mesocarp of coconuts |
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-simple fruit, hard exocarp, & fleshy mesocarp -ex: loofah & melons |
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-fleshy components of fruit do not develop from ovary -1 type: pome |
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-simple, accessory fruit, receptacle swells around ovary -ex: apple & pear -apple: fleshy components come from receptacle |
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-open to release seeds -classified based on how many places they open to release seeds -3 types: follicle, legume, & capsules |
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-dehiscent -simple, dry fruit that opens at 1 place -ex: milkweed & magnolia |
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-dehiscent -opens at 2 points to release seeds -ex: green beans & peanuts |
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-dehiscent -simple, dry fruits that open at more than 2 places to release seeds -ex: poppy |
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-4 types: achenes, samara, caryopsis, & nut |
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-indehiscent -dry fruits, seed attached to fruit wall at 1 point -ex: sunflower seed |
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-indehiscent -winged achene -ex: maple trees & dandelions |
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-indehiscent -produced by grass, pericarp is very thin & is fused to the seed coat -ex: grains from grasses |
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-indehiscent -similar to achene, but very strong pericarp |
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-produced from a flower w/many pistils in 1 flower -made-up of many fruitlets or drupelets -ex: blackberries & strawberries -strawberries: aggregate & accessory fruit, from receptacle, & has fruitlets |
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-many flowers grow close together -1 ovary is produced in each flower -fruits from multiple flowers grow close together & appear to fuse together -ex: pineapple |
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-closely related -both have vessel elements in xylem -both produce sporophylls that look like anthers -225 million yrs old |
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-oldest known angiosperm genus -130 million yrs old -means "old fruit" -seeds enclosed in carpels or fruits -fossils of fish close by, so possibly aquatic -no seeds or petals present |
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-Magnolia -possesses what's thought to be a primitive floral structure -possesses tepals: no differentiation between sepals & petals -lg numbers of organs in each whorl -spiral arrangement of organs |
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-"sister" to all flowering plants -in New Caledonia -sm flowers |
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Trends in Floral Evolution |
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-fewer organs in each floral whorls -4 whorls reduced to 3, 2, 1 -fusion of organs -superior to inferior ovary -radial to bilateral symmetry -sexual reproduction drives floral evolution -animal pollinated: may visit flower for reward (pollen & nectar) -Pollination Syndrome |
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-flowers w/traits preferred by specific organisms |
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Beetle Pollinated Flowers |
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-broad flowers -fruity-smelling |
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-may resemble rotting meat -fleshy & red -generate heat -smell putrid |
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-live on flower nectar & pollen -flowers usually brightly colored (purples, blues, yellows) -structures called "honey guides" that direct bee where to go to find the pollen -ultraviolet pigments |
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Moth & Butterfly Pollinated Flowers |
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-may have landing platform -usually white or pale-colored -sweet scent -spur |
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-long, hollow tube made of fused petals or sepals -may have nectar at base |
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-usually odorless b/c birds can't smell -flowers tend to be red |
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-copious nectar & pollen -dull color -musky scent -open at night |
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-animal (usually bee) doesn't visit flower for food reward -bee copulates w/flower that resembles another bee of same species |
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-catkin -grasses: wind-pollination |
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-sunflower family -flowers sm & packed together into a head -ray flowers & disk flowers |
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-both in family Asteraceae -ray: petals fused together into 1 long strap -disk: petals are fused into a cone around pistil, many fruits from 1 inflorescence (1 seed/fruit) |
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-lg.est angiosperm family -24,000 + species -massive fusion between & among all whorls -1,000's of ovules per ovary -genus: Vanilla (orchid) |
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-rose family -genus Malus (apples: pomes) -genus Pyrus (pears: pomes) -genus Rosa (roses) -genus Prunus (cherries, peaches, apricots: drupes) |
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-grass family -may be most economically important angiosperm family -growth form: rhizomes or stolons -leaves: linear w/parallel veins -flowers: very reduced, zygomorphic, wind-pollinated, sometimes arranged in spikes -fruit: caryopsis (grain) |
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-genus & species in Poaceae -cultivated rice -domesticated in Asia -lg. fruits selected for |
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-genus in Poaceae -wheat -most cultivated plant worldwide -used for bread production -protein: gluten |
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-pea family -herbs, shrubs, trees -leaves: usually compound -roots: produced nodules -flowers: usually zygomorphic -fruit: legume |
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-chambers that maintain an oxygen free envelope in which nitrogen-fixing bacteria can live |
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-violet family -flowers: zygomorphic -herbs or trees -2 types of flowers: Chasmogamous & Cleistogamous |
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-open flower that facilitates cross pollination |
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-closed flower -facilitates self-pollination -promotes inbreeding |
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-oak or beech family -temperate to tropical ecosystems in N Hemisphere -trees or shrubs -fruits: nuts (acorns) |
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-night shade family -herbs and trees -flowers: usually actinomorphic |
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-genus in Solanaceae -tobacco |
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-genus in Solanaceae -hot & bell peppers |
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-genus & species in Solanaceae -tomato |
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-genus & species in Solanaceae -potato |
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-mint family -growth form: usually herbaceous, secondary growth may be produced -stem morphology: usually square in cross section -leaf arrangement: opposite decussate -flowers: zygomorphic, corolla fused into tube -stamens: 2 lengths (2 sm. & 2 lg.) -volatile oils: oils evaporate into atmosphere, usually smell nice to us, classified as secondary metabolites, & may be used for defense |
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Trends in Plant Evolution: Bryophytes |
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-no vascular tissue -gametophyte is the dominant phase in life cycle -grown in moist, humid environments -spores for dispersal |
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Trends in Plant Evolution: Ferns & Vascular Non-Seed Plants |
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-Lycopodiophyta & Pteridophyta -branched sporophyte (dominant stage in life cycle) -gametophytes live in moist environment -water required for sexual reproduction |
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Trends in Plant Evolution: Seed Plants (Gymnosperms) |
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-seed is unit of dispersal -seed produced after sexual reproduction -seed contains embryo |
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Trends in Plant Evolution: Angiosperms |
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-flowers & fruits -help seed dispersal |
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