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All organisms share many characteristics: |
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Definition
•Composed of one or more cells •Carry out metabolism •Transfer energy with ATP •Encode hereditary information in DNA |
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Reconstruction and study of evolutionary relationships |
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Represents a hypothesis about patterns of relationship among species |
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that all species were descended from a single common ancestor He depicted this history of life as a branching tree “Descent with modification” |
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A derived characteristic is |
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Similarity that is inherited from the most recent common ancestor of an entire group |
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An ancestral characteristic is |
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Similarity that arose prior to the common ancestor of the group |
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Characters can be any aspect of the phenotype |
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Definition
Morphology Behavior Physiology DNA |
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exist in recognizable character states Example: Character “teeth” in amniote vertebrates has two states, present in most mammals and reptiles, and absent in birds and turtles |
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Presence of hair in mammals is ancestral or derived? |
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presence of lungs in mammals is ancestral or derived? |
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Diagram that depicts a phylogeny (hypothesis of evolutionary relationships) |
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Species that share a common ancestor as indicated by the possession of shared derived characters
Evolutionary units that refer to a common ancestor and all descendants |
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derived character shared by clade members |
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a shared character state that has not been inherited from a common ancestor |
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Systematists increasingly use |
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DNA sequence data to construct phylogenies Large number of characters can be obtained through sequencing
Character states are polarized by referencing the sequence of an outgroup |
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Assumes that the rate of evolution of a molecule is constant through time |
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How we place species and higher groups into the taxonomic hierarchy Genus, family, class, etc. |
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Includes the most recent common ancestor of the group and all of its descendants (clade) |
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Includes the most recent common ancestor of the group, but not all its descendants |
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Polyphyletic group does not include |
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the most recent common ancestor of all members of the group |
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Taxonomic hierarchies are based on |
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shared traits, should reflect evolutionary relationships |
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Biological species concept (BSC) |
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Definition
Defines species as groups of interbreeding populations that are reproductively isolated |
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Phylogenetic species concept (PSC) |
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Species is a population or set of populations characterized by one or more shared derived characters |
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PSC looks to the past for what? |
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to see if they have been separated long enough to develop their own derived characters |
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BSC is strictly _______ and PSC is _____ and _____ |
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sexual; asexual and sexual |
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The science of classifying living things |
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Shifted from emphasis on identifying and naming organisms to constructing evolutionary hypotheses to explain the relatedness of species
Organisms grouped into eight levels Domain, kingdom, phyl um, class, order, family, genus, and species Genus level and higher are capitalized Other categories assist with classification |
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Each hierarchal level is called a |
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Organisms grouped into eight levels |
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Domain, kingdom, phylum, class, order, family, genus, and species |
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What is the largest taxon |
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What is the most abundant organism on earth |
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Bacteria plays critical roles throughout the biosphere by |
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Carbon and sulfur cycling Extract nitrogen from air Photosynthesis 12 to 15 major groups of bacteria Based on ribosomal RNA sequences |
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Typically live in extreme environments Diverged early from bacteria More closely related to eukaryotes than to bacteria Based on genes that encode ribosomal RNAs Swapped genetic information via horizontal gene transfer (HGT) with other microorganisms |
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All archaea share certain key characteristics Lack peptidoglycan in cell walls Distinct lipid membrane structure Distinct rRNA sequences Genes have introns Grouped into three general categories Methanogens Extremophiles Nonextreme archaea |
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Most common model shows Eukarya branched from within Archaea Their structure and function allowed multicellular life to evolve The roots of the eukaryotic tree remain elusive |
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Eukarya molecular data support five monophyletic supergroups |
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Excavata S A R Archaeplastida Amoebozoa Opisthokonta |
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Eukaryotes have an extensive endomembrane system that |
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Divides cell into functional compartments |
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Mitochondria and chloroplasts entered early eukaryotic cells by |
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Compartmentalization of cells enabled the advent of |
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Bacteria and archaea are distinct from eukaryotes in that they have little or |
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– cells are differentiated, in contact and their functions are coordinated |
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Eukaryotic species as a group carry out _____ reproduction |
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First eukaryotes were probably ______ Diploids arose on separate occasions through ______ |
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Smallest and simplest, and most abundant forms of life |
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Prokaryotes likely originated on earth over_____ years ago Our knowledge of early life comes from microfossils Fall into 2 domains Bacteria Early cyanobacteria produced oxygen which allowed for diversity Archaea Earliest Archaea discovered were extremophiles |
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Our knowledge of early life comes from microfossils Fall into 2 domains Bacteria Early cyanobacteria produced oxygen which allowed for diversity Archaea Earliest Archaea discovered were extremophiles |
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Prokaryotes fall into 2 domains |
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Archaea resembles ______ instead of bacteria |
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three domains of life tree is based on ______ sequences |
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Prokaryotes are fundamentally different from eukaryote |
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bc most are single-celled, may stick together to form associations and biofilms Size varies tremendously Most are less than 1 µm in diameter
Single circular chromosome in nucleoid region Often have plasmids |
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Most prokaryote divide by_______, a form of asexual reproduction |
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Occurs through horizontal gene transfer Not a form of sexual reproduction |
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Single protein fiber Bacterial flagella rotate like propellers |
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Internal compartmentalization |
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Definition
No extensive membrane-bounded organelles Some have structures that resemble organelles Surrounded by a lipid membrane or protein shell Plasma membrane can be invaginated Isolates respiration or photosynthesis |
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Archaea and bacteria both lack a |
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Definition
membrane-bound nucleus and endomembrane system |
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archaea and bacteria have |
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Definition
distinctly different cell types. |
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Fatty acid chains joined to glycerol phosphate with ester linkages Stereoisomer of glycerol-3-phosphate Membranes are unbranched and are diethers |
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Non-polar hydrocarbons called isoprenoids joined to glycerol phosphate with ether linkages Stereoisomer of glycerol-1-phosphate Membranes can be branched, in cyclic compounds, and may be tetraethers that can form a monolayer membrane |
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Bacteria have _______ Archaea lack _____________ |
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S-layer outside of wall or replacing wall |
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Composed of glycoproteins that form a rigid paracrystalline surface |
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Evolved independently from bacterial flagella |
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bacteria and archaea Both have ______replication origin |
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All organically produced methane comes from archaea |
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only found in archaea and is a major contributor to global nitrogen cycle |
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Bacteria have 3 basic shapes |
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Definition
Bacillus – Rod-shaped Coccus – Spherical Spirillum – Helical-shaped |
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Peptidoglycan forms a rigid network |
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Unique to bacteria Maintains shape Withstands hypotonic environments |
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Gram-positive bacteria have a _______ peptidoglycan wall and stain a _______color |
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Gram-negative bacteria contain _______ peptidoglycan and (do not/do) retain the purple color |
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Gelatinous layer found in some bacteria Surrounds the cell wall Aids in attachment Protects from the immune system |
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Slender, rigid, helical structures Composed of the protein flagellin Involved in locomotion – spins like propeller |
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Short, hairlike structures Found in gram-negative bacteria Aid in attachment and conjugation |
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Endospores develop a thick wall around their _______ and a small portion of the _________ |
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Endospores form when exposed to |
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highly resistant to environmental stress Especially heat |
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Definition
tetanus, botulism, and anthrax |
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Bacterial microcompartments (BMCs) |
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Bound by semipermeable protein shell Function as storage or to isolate metabolic pathways |
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Prokaryotic cells often have complex ______ membranes |
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Prokaryotes do not reproduce sexually, there are 3 types of horizontal gene transfer |
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Conjugation – mediated by plasmids Transduction – mediated by viruses Transformation – direct uptake of DNA All 3 processes also observed in archaea |
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Virtually any gene can be transferred Occurs via |
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Definition
accidents in the lytic cycle |
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Term
When viruses go on to infect another cell |
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Definition
they transfer the bacterial DNA and it can be incorporated by homologous recombination |
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Specialized Transduction occurs via |
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Definition
accidents in the lysogenic cycle |
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DNA fragments are released from damaged or dead cells and are taken up by |
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The new DNA is incorporated into the live cell's genome by |
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The proteins needed for natural transformation are encoded by |
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Artificial transformation |
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Some species do not naturally undergo transformation Can be accomplished in the lab Critical for gene cloning and DNA manipulation |
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The ________ a gene is from the origin of transfer, the ________it will be transferred |
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energy from sun, convert inorganic C O2 to organic carbon |
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Definition
use H 2 S as electron donor, anoxygenic |
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energy from oxidizing inorganic substances |
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light as energy source but obtain organic carbon made by other organisms |
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both carbon atoms and energy from organic molecules |
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Definition
Second only to HIV/AIDS as cause of death by an infectious agent Agent: Mycobacterium tuberculosis Afflicts the respiratory system Easily transferred from person to person through the air In 2017, 10 million people contracted TB and about 1.6 millions died from the disease |
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Definition
Helicobacter pylori is the main cause Treated with antibiotics |
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Term
Dental caries (tooth decay) |
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Definition
Plaque consists of bacterial biofilms Streptococcus sobrinus and S. mutans ferments sugar to lactic acid Tooth enamel degenerates, leaving tooth vulnerable to infection |
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Term
Viruses or not organisms the have |
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Definition
No cellular structure, independent metabolism or reproduction Not alive or dead but inactive or active |
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Term
Most viruses come in two simple shapes |
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Definition
Helical - rodlike EX. Tobacco mosaic virus Icosahedral – soccer ball shape |
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Term
Some viruses are complex such as |
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Definition
Icosahedral head with helical tail Bacteriophages |
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Definition
Pithovirus virus Revived from 30,000-year-old Siberian ice in 2014 Virions are 1.5 µm long, larger than some bacteria |
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Definition
Production and release of new viruses Virus usually causes the cells to rupture, or lyse Allows infection of new cells or horizontal transmission of viruses |
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Latent phase Vertical transmission of viruses through cell division |
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Term
In lysogenic phage, expression of the viral genome is repressed by |
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Definition
a viral regulatory protein |
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Term
If the lysogen becomes stressed |
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Definition
the genome can be de-repressed |
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Term
Conditions for Pandemics Three conditions are necessary for a pandemic |
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Definition
New strain must contain a novel combination of H and N spikes
New strain must be able to replicate in humans and cause death New strain must be efficiently transmitted between humans. |
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Term
Eukaryotic cells differ from prokaryotes bc of |
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Definition
Presence of a cytoskeleton Compartmentalization (nucleus and organelles) |
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Term
The nucleus and endoplasmic reticulum arose from |
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Definition
infoldings of prokaryotic cell membrane |
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Many organelles evolved via endosymbiosis between an __________ and a ____________ |
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Definition
ancestral eukaryote and bacterial cell |
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Definition
Aerobic bacteria engulfed by larger bacteria |
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Larger bacteria engulfed smaller photosynthetic bacteria Chloroplasts come from single line of cyanobacteria Hosts are not monophyletic |
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engulfed red algae that already had chloroplasts |
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Endosymbiosis supported by |
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Definition
DNA inside mitochondria and chloroplasts DNA similar to bacteria DNA in size and character |
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Definition
Most diverse group of eukaryotes |
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Definition
Plasma membrane Extracellular material (ECM) in some Diatoms – Silica shells |
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Dormant cell with resistant outer covering Used for disease transmission |
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locomotion occurs how many time in flagella |
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Definition
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locomotion occurs how many time in cilia |
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Definition
shorter or more numerous than flagella |
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Definition
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Heterotrophic or autotrophic |
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Phagotrophs – Ingest particulate food matter |
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Photosynthetic Chemoautotrophic |
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both phototrophic and heterotrophic |
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Definition
From single cells to colonies to true multicellularity |
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Typical mode of reproduction Some species have an unusual mitosis |
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Definition
equal size daughter cells |
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one daughter cell smaller |
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cell division preceded by several nuclear divisions; produces several individuals |
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Definition
May be obligate, or only under stress |
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The Protists as a group are not |
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Definition
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Definition
This group consists of diplomonads, parabasalids, and euglenozoans |
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Unicellular Move with flagella 2 nuclei Giardia intestinalis- human parasite Mitochondrion-related organelles (MROs) |
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Live in termite guts Host cellulose degrading bacteria Trichomonas vaginalis – STD Undulating membrane for locomotion Use flagella Lack mitochondria, but have MROs – derived trait |
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Definition
Among the earliest eukaryotes to possess mitochondria One third have chloroplasts and are autotrophic Others lack chloroplasts and are heterotrophic All have a flexible pellicle No sexual reproduction |
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Two anterior (and unequal) flagella Attached at reservoir Contractile vacuoles – collect excess water Stigma – movement towards light Numerous small chloroplasts Symbiosis from ingestion of green algae |
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Second major group in Euglenozoa Unique, single mitochondrion DNA maxicircles and minicircles Trypanosomes cause human diseases African sleeping sickness – tsetse fly Leishmaniasis – sand fly Chagas disease – feces of triatomine bugs |
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Difficult to control because organisms repeatedly change their protective coat Release of sterilized flies Traps scented like cows but treated with insecticides Sequencing of genomes revealed core of common genes in all 3 – hope for single drug target |
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SAR: Stramenopiles and Alveolates |
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Definition
This is a supergroup consisting of three branches: Alveolata, Stramenopila, and Rhizaria |
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Brown algae, diatoms, and oomycetes Very fine hairs on their flagella A few species have lost their hairs during evolution |
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Conspicuous seaweeds of northern regions Life cycle involves alternation of generations Sporophyte – multicellular and diploid Gametophyte – multicellular and haploid Not plants |
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Phylum Chrysophyta Photosynthetic, unicellular organisms Unique double shells made of silica Some move using raphes Two long grooves lined with vibrating fibrils |
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“Water molds” Either parasites or saprobes Were once considered fungi Motile zoospores with two unequal flagella Produced asexually Undergo sexual reproduction Found in water or on land Phytophthora infestans Irish potato famine (1845 to 1847) |
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Definition
Flattened vesicles called alveoli Common lineage despite diverse modes of locomotion |
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three groups of alveolata |
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Definition
Dinoflagellates Apicomplexans Ciliates |
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Definition
Photosynthetic, unicellular with two flagella Live in aquatic environments Some are luminous Do not appear to be directly related to any other phylum “Red tide” are “blooms” – fish, birds, and marine mammals may die from toxins DNA not complexed with histones |
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Definition
unique arrangement of organelles at one end of the cell which enables the cell to invade its host
Most common in Africa Control focused on vector control and insecticide-treated mosquito nets Resistance to treatment and insecticides continues to be a problem Vaccine, Mosquirix, only moderately effective |
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Definition
Found in the intestines of arthropods, annelids, and mollusks |
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Definition
Causes infections in humans with immunosuppression Can cross placental barrier to harm fetus |
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Definition
tough but flexible outer covering |
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Definition
Micronucleus – without will reproduce asexually Macronucleus – essential for function |
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Have two types of vacuoles |
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Food vacuoles – digestion of food Contractile vacuoles – regulation of water balance |
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Definition
Rhizaria use pseudopods for locomotion |
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Three distinct monophyletic groups of Rhizaria |
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Definition
Radiolaria, Foraminifera, and Cercozoa |
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Actinopoda (Radiolarians) |
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Definition
Glassy exoskeletons made of silica Needlelike pseudopods |
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Definition
Heterotrophic marine protists Pore-studded shells called tests, through which thin podia emerge Use podia for swimming and feeding Complex life cycles with haploid and diploid generations |
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Definition
A morphologically diverse group of primarily soil protists Locomotion with flagella or pseudopods Some have silica-based shells made of scales or plates |
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Definition
consists of Rhodophyta, Chlorophyta, Charophytes, and land plants |
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Definition
Red algae range from microscopic to very large Lack flagella and centrioles Have accessory photosynthetic pigments within phycobilisomes Origin has been a source of controversy Tentatively, treated as a sister clade of Chlorophyta (green algae) |
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Definition
Land plants arose from an ancestral green alga only once during evolution |
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Green alga consist of 2 monophyletic groups |
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Definition
Chlorophyta Streptophytes (contains the Charophytes) |
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Definition
Early green algae probably resembled Chlamydomonas reinhardtii Diverged from land plants over 1 BYA Several lines of evolutionary specialization derived from chlorophytes |
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Multicellular Chlorophytes |
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Definition
Cell specialization in colonial chlorophytes Multicellularity arose many times in the eukaryotes Colonial chlorophytes are examples of cellular specialization |
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Term
Haplodiplontic Life Cycles in Multicellular Chlorophytes |
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Definition
Ulva Multicellular chlorophyte Identical gametophyte and sporophyte generations Consist of flattened sheets two cells thick |
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Term
Multicellular diploid stage |
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Definition
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Term
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Definition
Produces haploid spores by meiosis Diploid spore mother cells (sporocytes) undergo meiosis in sporangia Produce 4 haploid spores First cells of gametophyte generation |
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Term
Multicellular haploid stage |
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Definition
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Term
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Definition
Spores divide by mitosis Produces gametes by mitosis Gametes fuse to form diploid zygote First cell of next sporophyte generation |
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Charophytes distinguished from chlorophytes by |
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Definition
phylogenetic relationship to land plants |
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Definition
haplontic life cycles Evolution of diplontic embryo and haplodiplontic life cycle occurred after move to land |
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2 candidate Charophyta clades |
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Definition
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Both charophyte clades form _________ around the edges of freshwater ponds and marshes One species must have successfully inched its way onto land through adaptations to drying |
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Definition
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Term
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Definition
Amoebas move by means of pseudopods |
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Definition
flowing projections of cytoplasm Extend and pull the amoeba forward Engulf food particles |
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Definition
free living Found in the soil as well as freshwater Some are parasitic Acanthomoeba enters the body through a wound and crosses the blood-brain barrier into the brain |
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Definition
Were once considered fungi |
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Definition
include two lineages Plasmodial slime molds Huge, single-celled, multinucleate, oozing masses Cellular slime molds Single cells combine and differentiate, creating an early model of multicellularity |
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Definition
Stream along as a plasmodium Nonwalled, multinucleate mass of cytoplasm Form called feeding phase Ingests bacteria and other organic material When food or moisture is scarce, organism forms sporangia, where spores are produced |
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Definition
Important group for the study of cell differentiation because of their relatively simple developmental systems |
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Definition
Colonial choanoflagellates resemble their close animal relatives, the Sponges Choanoflagellida are likely the ancestors of animals |
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Term
Mycologists believe there may be as many as _______ fungal species |
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Definition
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Term
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Definition
Single-celled or multicellular Sexual or asexual Specialized to extract and absorb nutrients from surroundings Animal and fungi share several characteristics |
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Definition
Multicellular fungi consist of long, slender filaments |
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Term
Some hyphae are continuous called ___________hyphae Others are divided by |
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Definition
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______flows throughout hyphae |
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Definition
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Term
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Definition
mass of connected hyphae Grows through and digests its substrate |
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Term
Fungal cell walls include |
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Definition
chitin Also found in the hard shells (exoskeletons) of arthropods Both fungi and animals store glucose as glycogen |
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Term
Hyphae may have more than one nucleus |
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Definition
Monokaryotic – 1 nucleus Dikaryotic – 2 nuclei |
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Term
Fungi have an unusual mitosis |
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Definition
Cell is not relevant unit of reproduction Nuclear envelope does not break down and re-form Instead, the spindle apparatus is formed within it |
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Term
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Definition
centrioles (except in chytrids) |
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Term
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Definition
Capable of both sexual and asexual reproduction |
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Term
Fungi Sexual reproduction |
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Definition
Fusion of two haploid hyphae of compatible mating types In some fungi, fusion immediately results in a diploid (2n) cell Others, have a dikaryon stage (1n + 1n) before parental nuclei form diploid nucleus |
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Term
______ are the most common means of reproduction among fungi May form from sexual or asexual processes Most are dispersed by wind or insects |
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Definition
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Term
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Definition
Obtain food by secreting digestive enzymes into surroundings Then absorb the organic molecules produced by this external digestion Great surface area-to-volume ratio |
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Definition
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Term
Fungi, together with bacteria, |
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Definition
are the principal decomposers in the biosphere |
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Definition
essential for fungus survival |
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Definition
nonessential for survival |
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Definition
pathogens harm host by causing disease |
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Definition
benefit one partner but does not harm the other |
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Term
Mutualistic relationships |
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Definition
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Term
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Definition
cause harm to host (do not cause disease) |
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Term
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Definition
Live in the intercellular spaces inside plants Some parasitic, some commensalistic Some fungi protect their hosts from herbivores by producing toxins |
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Term
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Definition
Symbiotic associations between a fungus and a photosynthetic partner |
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Term
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Definition
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Definition
unable to grow normally without their photosynthetic partners |
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Term
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Definition
Mutualistic relationships between fungi and plants Found on the roots of about 90% of all known vascular plant species Function as extensions of root system Increase soil contact and absorption of nitrogen, phosphorus and race elements |
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Term
Two principal types of mycorrhizae |
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Definition
Arbuscular mycorrhizae Ectomycorrhizae |
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By far the most common Fungal partners are glomeromycetes No aboveground fruiting structures Possibly helped early vascular plants survive infertile soils |
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Most hosts are forest trees (pines, oaks) Fungal partners are mostly basidiomycetes |
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Ruminant animals host fungi in their gut that can degrade cellulose and lignin Leaf-cutter ants have domesticated fungi which they keep in underground gardens Ants provide fungi with leaves Fungi are food for the ants |
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Fungi may secrete substances making food unpalatable, carcinogenic, or poisonous for example |
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Aspergillus flavus – aflatoxin Ustilago maydis – edible corn smut (not harmful to animals) |
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Fungi also cause human and animal diseases such as |
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Definition
Candida – thrush; vaginal infections Ringworm, athlete’s foot, and nail fungus |
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Responsible for the worldwide decline in amphibian populations |
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Batrachochytrium dendrobatidis causes |
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Microsporidia Blastocladiomycota Neocallimastigomycota Chytridiomycota Glomeromycota Basidiomycota Ascomycota |
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1 phylum that is not monophyletic |
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Definition
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Obligate, intracellular, animal parasites Long thought to be protists Lack mitochondria Have reduced mitochondrion-like organelles |
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Polar Tube of Microsporidians |
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Definition
Commonly cause disease in immunosuppressed patients Infect hosts with their spores, which contain a polar tube |
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Aquatic, flagellated fungi Closely related to ancestral fungi Have motile zoospores |
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Definition
Alternation of generations- only fungi that do so |
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Definition
Grows in the soil Both diploid and haploid stages are multicellular |
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Spore producing via meiosis for haploid spores Mitosis used to produce diploid spores for asexual cycle |
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Definition
gamete producing via mitosis |
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Digest plant biomass in mammalian herbivore rumens Mammal depends on fungi for sufficient calories Lack true mitochondria Horizontal gene transfer brought cellulase gene from bacteria into Neocallimastix genome Potential use for biofuel production- degrade cellulose at high temperatures |
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are not monophyletic – still under research Include the common bread molds, a few human pathogens Mostly coenocytic hyphae Chemotropic hyphae are attracted to opposite type in unfavorable conditions for sexual reproduction cycle |
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Life Cycle of Zygomycetes Sexual reproduction |
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Definition
Fusion of gametangia Haploid nuclei fuse to form diploid zygote nuclei – karyogamy Develops into zygosporangium in which zygospore develops Meiosis occurs during germination of zygospore Releases haploid spores |
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Asexual reproduction of zygomycetes |
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Definition
more common Sporangiophores have sporangia that release spores |
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Definition
tiny group of fungi Form intracellular associations with plant roots called arbuscular mycorrhizae Cannot survive in absence of host plant No evidence of sexual reproduction Likely instrumental in colonization of land by plants |
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are some of the most familiar fungi Mushrooms, toadstools, puffballs, shelf fungi, etc. Also important plant pathogens like rusts and smuts |
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club-shaped sexual reproductive structure |
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Spore germination leads to |
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the production of monokaryotic hyphae Results in a monokaryotic mycelium, or primary mycelium |
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Different mating types of monokaryotic hyphae may fuse Results in a |
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Definition
dikaryotic mycelium, or secondary mycelium |
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Contain about 75% of the known fungi Includes bread yeasts, common molds, cup fungi, truffles, and morels Serious plant pathogens – cause of chestnut blight and Dutch elm disease |
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producing fungi are in the genus Penicillium |
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Conidia formed at the ends of modified hyphae called |
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Definition
conidiophores Allow for the rapid colonization of a new food source Many conidia are multinucleate |
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Ascomycetes named for ascus – |
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Definition
microscopic, saclike reproductive structure |
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Unicellular ascomycetes Most reproduce asexually by budding Yeasts can ferment carbohydrates Break down glucose into ethanol and C O2 Used to make bread, beer, and wine Saccharomyces cerevisiae |
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Yeast can ferment simple sugars Likely a response to |
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Definition
competition with other microorganisms (bacteria) |
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Yeast can ferment sugar in the absence of oxygen having an advantage |
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Definition
over those organisms that need oxygen as the final electron acceptor |
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