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Theory of Uniformitarianism
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By James Hutton
Geological processes today also occured in the past
we observe the present to learn about the past |
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study of primate/human evolution |
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before present
years ago
thousands of years ago
millions of years ago |
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relative dating method
determines place of object in sequence of time, not exact |
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study of rock strata and the chronology of those layers |
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layers on top are newer, bottom are older |
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relative
accumulation of flouride levels in bone
no between site comparisons
the higher the flouride count, the older the sample |
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absolute dating method
radioactive
used w/ organic matter
half-life= 5,730 years
how long it takes Carbon-14 to turn into Nitrogen-14
one neutron changes into a proton |
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used with volcanic rocks older than 100,000 years
Half-life=1.25 billion years
dates the strata not the fossil
40K -> 40Ar
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Better than K-Ar dating
find proportion of 39Ar to 40Ar
one step process
faster, more accurate |
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Sexual dimorphism (pelvis, cranium)
age (dental eruption, epiphysial fusion) |
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focus on similarities and lump specimens into existing genera |
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classify specimens into new species |
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study of what happens to an organism after it dies |
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study of what's happened to the environment |
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hands-on: attempts to make the stone tools, etc. |
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What determines units of time? |
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Geologic, climatic and biological events |
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Precambrian Large Chunk of Time |
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Composed of Hadean, Archean, and Proterozoic Eons
About 90% of Earth's history |
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origin of Earth 4.6 billion years ago-> 542 million years ago |
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single-celled organisms
1st multi-celled organisms |
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Cambrian Large Chunk of Time |
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1 Eon= Phanerozoic
Current Large Chunk of Time |
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Part of Phanerozoic Eon
542-251mya
"The Cambrian Explosion!!"- lots of speciation
origin of 1st vertebrates and fish
Tiktaalik- transitional species from fish to land
dinosaurs and birds
Therapsids- mammal-like reptiles |
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Permian-Triassic Mass Extinction |
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Hydrogen sulfide
Volcanic eruptions-> global warming-> bacteria in oceans-> hydrogen sulfide emissions-> Mass extinction of 75% terrestrial and 95% marine life
-OR-
Mantle superplume-> weakened magnetic field-> cosmic radiation-> cloud coverage-> cooling-> death-> Permian-Triassic volcanic eruption
(Yukio Isozaki) |
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241-65.5mya
"Age of Dinosaurs"
200mya- true mammals
(monotremes
marsupials
placentals) |
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3 periods w/in Mesozoic Era |
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Triassic
Jurassic
Cretaceous |
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65mya-present day
"Age of Mammals"
primate and human evolution |
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Paleocene
Eocene
Oligocene
Miocene
Pliocene
Pleistocene
Holocene |
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Late Ordovician
Late Devonian
Permian-Triassic
Late Triassic
Cretaceous-Tertiary |
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Pangea-> Laurasia-> Gondwana |
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long-term evolutionary chance |
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65.5mya-55.8mya adaptive radition of plesiadapiforms |
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arboreal small long snout lateral orbits lack of postorbital bar large diastema forward positioned anterior teeth claws |
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56-34mya adaptive radiation of 1st true primates/early prosimians adapidae and omomydae |
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reduced snout forward facing orbits grasping hands reduced snout Diurnal folivores and frugivores broadly similar to modern lorises and lemurs |
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reduced snout forward facing orbits presence of postorbital bar teeth closer together smaller, nocturnal frugivores and insectivores broadly similar to modern tarsiers haplorrhine? |
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Models of Primate Origins |
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3 main models-> grasping hands and stereoscopic vision 1-arboreal lifestyle: evolve @ same time 2- vesual predation: adapts for hunting insects 3- terminal branch feeding- grasping hands developed for gathering fruits and flowers for eating |
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type of plesiadapiform no stereoscopic vision, but DOES have grasping hands (thus, hands evolved before eyes) intermediate morphology between primate-like mammals and true primates |
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33.9-23mya adaptive radiation of 1st anthropoids continued reduction of snout- more reliance on vision enclosed orbits small, diurnal, arboreal, typically quadrupeds,mainly frugivorous |
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single origin of anthropoids in Africa 30mya "rafting" to South America |
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23-5.3mya adaptive radiation of 1st hominoids hominin origins great diversity in ape species |
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mix of monkey and ape traits quadruped larger cranial capacity relative to body size no tail arboreal frugivore |
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Early Hominin Cranial Capacity and Body Size |
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measure of the interior volume of the brain case measured in cubic centimeters average 400-500cc (ape size) modern human- 1350 cc average- 100lbs, 3.5 ft tall |
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Sahelanthropus Orrorin Ardipithecus |
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Sahelanthropus tchandensis |
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found in Chad Late Miocene (7-6mya) small, ape-like brain small canines foramen magnum position suggests bipedalism no post-cranial remains |
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Tugen Hills, Kenya late Miocene femur=bipedal humerus and phalanges= still partly arboreal |
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middle Awash, Ethiopia Pliocene (5.8-4.4mya) dentition=large, ape-like canines small brain size foramen magnum=bipedal primitive facial traits |
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5.3-1.8mya numerus species of hominins 1st homo sapiens |
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Australopithecus Anamensis |
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Kenya and Ethiopia 4.2-3.9mya distal tibia=bipedal large canines parallel dental arcade transitional form |
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Australopithecus afarensis |
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Hadar, Ethiopia (Don Johnson 1974) Laetoli, Tanzania (Mary Leakey 1974) 3.7-3mya small brain prognathic face femur+tibia=bipedal |
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Australopithecus afarensis Don Johnson female 40% of skeleton found 3'3, 60lbs fully erupted 3rd molars bipedal |
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Robust Austrolopiths/Paranthropus |
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2.5-1.4mya large jaws large molars large face promient sagittal crest |
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A. aethiopocus (east Africa) A. robustus (S. Africa) A. boisei (E. Africa) |
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Australopitecus africanus |
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3.3-2.5mya S. Africa lack of massive features possible ancestor to genus Homo |
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2.5mya Ethiopia similar features to A. afarensis possible ancestor to genus Homo |
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Ardipithecus ramidus -> A. anamensis-> A. afarensis -> A. africanus -> A. garhi -> Homo |
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1912, Piltdown England larger brain size? Flouride analysis Modern human skull and orangutan jaw messed up classification for a long while |
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Locomotion: Hominin-Ape Common Ancestor 3 possibilities |
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Bipedal Generalized pattern of locomotion Knuckle walker (wrist bones) |
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What came first? Bipedalism or Big Brains? |
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increased cranial capacity flatter face reduced dentition decreased brow ridges wider superior cranium and narrower base rounder occipital region forehead/vertical frontal bone presence of a chin |
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manufacture and use of stone tools |
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2-1.6mya: Pliopleistocene Olduvai Gorge, Tansania 1960 Lois and Mary Leakey earliest fossil evidence for Homo larger brain size smaller teeth still partly arboreal Oldowan tradition scavengers |
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Very similar to H. habilis. Different species or the same? |
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Homo erectus "Upright man" |
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1.8mya-800kya: Pleistocene Africa, E. Europe, and Asia Acheulian tradition More specialized tools for scraping and cutting scavenger/hunter FIRE!!! 1st Hominin species totally terrestrial modern patterns of bipedal locomotion first to move out of Africa Larger brain size occipital ridge sagittal keeling |
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Homo ergaster "Working man" |
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heat light scare off animals cook |
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H. heidelbergensis "Heidelberg man" |
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800-200kya: Pleistocene Europe, Africa, and Asia Brain size larger than H. erectus and almost as large as H. sapiens large face prominent brow ridges Levallois technique wooden spears hunter |
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Atapuerca, Spain 1.2mya-800kya Ancestor to or the same as H. heidelbergensis? |
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H. neanderthalensis "Neandertal man" |
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130-28kya: Pleistocene Europe and Middle East large brains large browridges lack of chin same vocal ability as modern humans occipital bun large nose short and stocky build (robust morphology) Mousterian tradition hunters buried dead care for sick and elderly stone tool technology |
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DNA evidence H. neanderthalensis |
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osteology and pathology reached older ages mtDNA b/w humans is very different suggests not ancestors to us nuclear DNA suggests split from common ancestor 500kya |
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H. floresiensis "the Hobbit" |
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95-12kya 2003 Flores, Indonesia 3.5 ft adults small cranial capacity mosaic pattern microcephalic H. sapien? |
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Why were H. floresiensis so small? 4 theories |
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1. process by which natural selection favors smaller body sizes on an island w/ less resources leading to dwarfism. 2. Heat and humidity could also lead to dwarfism- thermoregulatory condition 3. Genetic isolation/low gene flow 4. absence of terrestrial predators |
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Lee Burger 1-3kya Palau, Micronesia 2006/2007 similar body size as H. floresiensis, but larger cranial capacity; derived pelvic morphology another example of island dwarfism? island-adapted H. sapiens |
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Anatomically modern H. sapiens |
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Characteristics of A.M. H sapiens |
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well rounded skull wider superior cranium and narrower @ base reduced prognathism reduced brow ridges forehead vertical frontal bone presence of a chin |
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Lower paleolithic (2.5mya-300kya): stone tool technologies of H. habilis, H. rudolfensis, and H. erectus Middle Paleolithic (300-40kya) H. heidelbergensis and H. neanderthalensis Upper paleolithic (40-10kya) H. sapiens |
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natural: caves/rock shelters manufactured: huts- wood, animal, bone, animal hides |
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Lascout cave, France 30kya Europe, Africa, and Australia large game animals and hunting |
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past future: sympathetic Magic cultural symbolism art for art's sake |
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Shell beads bear teeth beads figurines fertility figures? other religious symbol? art? toys? |
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1.5mya erectus moves out of africa modern humans expand further 150kya humans move out of Africa reach Australia as early as 60kya and the New World 40kya |
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Transition to Modern Humans 3 models |
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Multiregional evolution: multiple areas of origin African replacement: started in Africa and spread w/o mixing w/ other species Assimilation: started in Africa and spread w/ interbreeding |
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the appearance of similar traits w/in a geographic region that remain over a long period of time |
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Europe: high-tented nose East Asia: shovel shaped incisors |
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African replacement model |
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most likely H. sapiens maybe technologially superior H. sapiens and Neanderthals maybe produced infertile offspring H. sapiens and Neanderthals did not mate due to social barriers |
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no Neanderthals contributed to modern gene pool greatest diversity in African populations mtDNA: 2 cluster both w/ African ancestry= most recent common ancestor African all allels found in non-African populations are found in African populations Max Planck Institute, Germany |
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Mechanism of Change Towards Modern Humans |
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Technology: increased complexity in tools-> food preparations-> less heavy chewing-> reduction in size of cranial features-> energy goes to increase brain size
Expensive tissue hypothesis: Aiello and Wheeler |
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