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an assemblage of populations of various species living close enough for potential interaction |
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Integrated Hypotheses (Clements) and Individualistic Hypotheses (Gleason) |
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A community is an assemblage of closely linked species, locked into association by mandatory biotic interactions -Predicts that the presence or absence of particular species depends on the presence or absence of other species -One should see sharp ecotones between distinct communities with little overlap in the species present in alternative communities |
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Individualistic Hypotheses |
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Proposes that communities are loosely organized associations of independently distributed species with the same abiotic requirements -Predicts that each species is distributed according to its tolerance ranges for abiotic factors |
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Populations in a community are potentially linked by |
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interspecific interactions -A community’s interactions include competition, predation, herbivory, symbiosis, and disease |
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Interspecific competition |
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occurs when species compete for a particular resource |
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Strong competition can lead to |
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competitive exclusion -the local elimination of one of the two competing species |
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Gause studied competition in laboratory experiments using |
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Paramecium species -Each species, when alone, shows logistic growth to a carrying capacity -When both species are present, one is driven to extinction by the other -Conclusion: two species cannot coexist if their niches are too similar |
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The Competitive Exclusion Principle |
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States that two species competing for the same limiting resources cannot coexist in the same place |
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The ecological niche is the multidimensional space that an organism occupies. The dimensions of the niche include abiotic factors as well as biotic resources and interactions -Two Types: Fundamental and Realized |
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is the space occupiable in the absence of biotic interactions |
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is the space the organism actually occupies |
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As a result of competition |
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a species' fundamental niche may be different from its realized niche |
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The Niche concept allows restatement of the |
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competitive exclusion principle -two species cannot coexist in a community if their niches are identical |
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Ecologically similar species can coexist in a community only if |
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there is one or more significant difference in their niches |
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the differentiation of niches that enables similar species to coexist in a community |
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-The tendency for characteristics to be more divergent in sympatric populations than in allopatric populations of the same two species -Evidence of divergence due to competition |
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predator and prey populations can strongly interact (eg. lynx and hare) -Predation shapes many attributes of both predators and prey: -Claws, teeth, fangs, stingers, and poison of predators -crypsis, mimicry, shells, speed, sensory capabilities of prey |
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Cryptic coloration, or camouflage makes prey |
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warns predators to stay away from prey |
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a palatable or harmless species mimics an unpalatable or harmful model -ex. hawkmoth larva, green parrot snake |
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two or more unpalatable species resemble each other |
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the process in which an herbivore eats parts of a plant -Has led to the evolution of plant mechanical and chemical defenses and consequent adaptations by herbivores |
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one organism, the parasite, derives its nourishment from another organism, its host, which is harmed in the process -ex Cymothoa exigua and the rose snapper; parasite replaces rose snappers tongue to share food |
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an interspecific interaction that benefits both species -Mutualisms are systems of mutual exploitation and can break down into interactions that benefit one player at the expense of the other |
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-tree provides food, nest sites for ants -ant provides tree protection from competitors, herbivores |
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one species benefits and the other is not affected |
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Two components of Species Diversity |
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-number of species -relative abundance of each species (evenness) |
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the feeding relationship between organisms in a community -is a key factor in community dynamics -food chains, food webs, or trophic pyramids |
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link the trophic levels from producers to top carnivores -usually only a few links long -in many studies, there is a correlation between productivity and food chain length |
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a branching food chain with complex trophic interactions |
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Certain species have an especially large impact on the structure of entire communities either because |
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they are highly abundant or because they play a pivotal role in community dynamics -dominant and keystone species |
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-Are those species in a community that are most abundant or have the highest biomass -Exert powerful control over the occurrence and distribution of other species |
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-Are not necessarily abundant in a community -Exert strong control on a community by their ecological roles, or niches |
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Ecosystem 'Engineers' (Foundation Species) |
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Some organisms exert their influence by causing physical changes in the environment that affect community structure |
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Some foundation species act as |
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facilitators -have positive effects on the survival and reproduction of some of the other species in the community |
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2 Models of control that communities can be subjected to |
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bottom-up and top-down controls, or both |
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-Proposes a unidirectional influence from lower to higher trophic levels -in this case, the presence or absence of abiotic nutrients determines community structure, including the abundance of primary producers |
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-Proposes that control comes from the trophic level above -The effects of top down control vary with the length of the food chain |
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-Is an event that changes a community -Removes organisms from a community -Alters resource availability -Fire: Is a significant disturbance in most terrestrial ecosystems and is often a necessity in some communities (helps eradicate dead grass) |
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The large-scale fire in Yellowstone National Park in 1988 |
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Demonstrated that communities can often respond very rapidly to a massive disturbance |
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orderly build-up of a community following disturbance and may proceed by any or a combination of tolerance, facilitation, and inhibition -Diversity might be highest at intermediate rates of disturbance |
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Occurs where no soil exists when succession begins -retreating glaciers provide a valuable field-research opportunity on this type of succession |
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Begins in an area where soil remains after a disturbance |
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three models of succession |
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Facilitation, Inhibition, Tolerance -Facilitation: Early arriving species may facilitate the appearance of later species by making the environment more favorable -Inhibition: Early arriving species may inhibit establishment of later species -Tolerance: Early arriving species have no impact on later arriving species |
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Intermediate disturbance hypothesis |
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-Suggests that moderate levels of disturbance can foster higher species diversity than low or high levels of disturbance -Frequently-disturbed habitats harbor primarily early successional species -Rarely-disturbed habitats harbor primarily climax species -Intermediate disturbance levels lead to highest levels of diversity |
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Two Key factors correlated with a community's species diversity |
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-geographic location (for instance latitude) -size |
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Species richness generally declines along an |
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along an equatorial-polar (latitudinal) gradient -Diversity is usually highest near the equator (tropics) and declines toward the poles -Many factors may contribute to this pattern including: Greater age of tropical environments (more time for the evolution of species) and Greater Productivity in the tropics due to greater sunlight and water availability |
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two main climatic factors correlated with biodiversity |
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temperature, solar energy input and water availability which are combined in measures of evapotranspiration -*correlation need not imply causation* |
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Latitudinal gradients in species richness may have both |
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historical and ecological determinants |
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Species-area curve quantifies the idea that |
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All other factors being equal, the larger the geographic area sampled, the greater the number of species |
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Equilibrium Model of Island Biogeography |
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Species richness on islands represents a balance between the rates of immigration and extinction -explains variation in the species diversity of islands through the effects of island size and distance from source populations. -Size of island affects both immigration and extinction rates • Larger islands have higher rates of immigration and lower rates of extinction -Distance from the source (mainland) affects immigration rates • Immigration more frequent if island nearer mainland -ex. Species richness increases with island size in the Galapagos and many other archipelagos |
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