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Hearing and Balance (Vestibulocochlear Organ) Hearing (Transformation of mechanical sound waves into nerve impulses) Balance (Changes in fluid levels within inner ear) |
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Parts of the External Ear |
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Auricle or Pinna - “Ear” External Acoustic Meatus - Canal connecting base of auricle to tympanic membrane (eardrum) |
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Funnel Shaped: Wide distally, Narrows to connection with external acoustic meatus Mobile: Can be orientated towards sound Superficial muscles |
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Determined by auricular cartilage Variable stiffness May maintain position of ear at all times |
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Cartilages of External Ears |
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Tympanic cavity (contains but often referred to as tympanic bulla) Tympanic Membrane Auditory Ossicles Auditory tube (Eustacian Tube) |
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Divides External Acoustic Meatus from Middle Ear Tilted Surface area greater than cross section of meatus Epidermal (outer) and mucosal (inner) covering Fibrous attachment to tympanic ring of temporal bone Medial surface attached to ossicles |
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Malleus (“Hammer”) Incus (“Anvil”) Stapes (“Stirrup”) Handle/Manubrium of malleus attached to eardrum Base of stapes inserts into vestibular window |
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Connects tympanic cavity to nasopharynx Openings in lateral wall of nasopharynx Runs in cartilage trough Out-pouching in horses = Guttural Pouch Usually collapsed When open allows equalisation of pressure on either side of tympanic membrane Allows drainage of material from tympanic cavity |
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Hearing - transformation of mechanical stimuli (sound) into nerve impulses Organs of balance |
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Maculae (otolith organs) Crystals (statoconia, otoliths) adhere to sensory hairs Determine position of head relative to gravity |
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Orientated at right angles Anterior, Posterior and Lateral Ampullae at one insertion into utriculus Fluid moves with head movement |
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Central Chamber: Vestibule (Contains ultriculus and sacculus) Semicircular Canals: Contain semicircular ducts Cochlea: Contains cochlear duct, Spiral – resembles snail shell, Central osseous pyramid (modiolus) with canal running around and projecting spiral lamina |
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Membrane from spiral lamina divides into three channels: Scala Vestibuli, Cochlear Duct, Scala Tympani |
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Vibrations at vestibular window (from stapes) and cochlear window Transmitted through perilymphatic space Causes vibration of basement membrane and movement of sensory cilia Registered by neuroreceptors Transmitted to vestibulocochlear nerve (VIII). |
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Nerve Supply to Inner Ear |
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Vestibulocochlear nerve CN VIII) Divides into vestibular and cochlear parts at internal acoustic meatus Vestibular – balance Cochlear – hearing Facial nerve (CN VII) Stapedius m Mandibular nerve (CN V) Tensor tympani m |
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Relationship of nerves in the face |
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Facial nerve passes through inner and middle ears Sympathetic supply to eye passes between tympanic bulla and petrous temporal bone Both can be damaged by ear disease |
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External Ear: Arterial: external carotid via caudal auricular Venous: maxillary vein Middle and Inner ear Arterial: Occipital Artery, branching from external carotid |
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Diagnostic Tools for the Ear |
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Otoscope: Direct examination of external ear Radiography: Widely available Tympanic bulla superimposed on other skull structures Advanced Imaging (CT/MRI): Clear imaging of inner and middle ear, Expensive |
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Inflammation or Infection of ear Otitis Externa: External Ear Otitis Media: Middle Ear Otitis Interna: Inner Ear |
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Embryological Development of the Ear |
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External Ear: Ectoderm of 1st Branchial Groove Middle Ear: Endoderm of 1st pharyngeal pouch Inner Ear: Ectoderm lateral to hindbrain |
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Hearing Range of Animals (pitch) |
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Humans: 20 – 20,000 Hz Dogs: 67 – 45,000 Cats: 45 – 64,000 Horse: 55 – 33,500 Elephant: 16 – 12,000 Chicken: 125 – 2000 Bat: 2000 – 110,000 |
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Cochlea converts sound (pressure waves) into action potentials |
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